Introduction. The name of each listed species is followed by up to five sources that report the species as occurring in Michigan. Sources in all upper-case letters refer to specific insect collections: MSU-ARC refers to material held by the Michigan State University Arthropod Research Collection (data provided by G. Parsons). All others refer to collection data extracted from the Symbiota Collections of Arthropods Network (SCAN) database online at https://scan-bugs.org/. See the Methods page for a list of abbreviations and associated collections.

Species names preceded by a single '?' are considered probable (for occurring in Michigan) but uncertain records. Those preceded by '??' are considered possible but unlikely to occur in Michigan. Species that are fully accepted as having valid Michigan records are unmarked. Species reported in the literature that we consider to be erroneous Michigan records are in a separate list following the others.

STAPHYLINIDAE - MISCELLANEOUS SMALLER SUBFAMILIES

12 subfamilies, 8 tribes, 35 genera, 93 species

Subfamily Euaesthetinae

Tribe Euaesthetini

Edaphus americanus Puthz, 1974: Herman (2001), Downie and Arnett (1996)

?Edaphus nitidus Motschulsky, 1857: Hubbard and Schwarz (1878), SI:NMNH; we assume H&S had authority wrong; if not, we don't know what this is

Euaesthetus albertae Puthz, 1998: Puthz (2014), Herman (2001), Casey (1884), Downie and Arnett (1996); syn. brevipennis described from Detroit by Casey (1884); BugGuide still uses brevipennis (in 2023)

Euaesthetus americanus Erichson, 1840: Herman (2001), Hubbard and Schwarz (1878), Leng (1920), Downie and Arnett (1996), MSU-ARC

Euaesthetus ganglbaueri Bernhauer, 1928: Herman (2001), Downie and Arnett (1996)

Euaesthetus iripennis Casey, 1884: Puthz (2014), Downie and Arnett (1996)

Euaesthetus mundulus Casey, 1884: Puthz (2014)

Euaesthetus pugetensis Hatch, 1957: Puthz (2014)

Euaesthetus suteri Puthz, 2014: Puthz (2014)

Subfamily Habrocerinae

Tribe Habrocerini

?Habrocerus capillaricornis (Gravenhorst, 1806): KU:SEMC; Wayne Co, Redford Township, Bell Creek Park

Habrocerus magnus LeConte, 1878: LeConte (1878), Wolcott (1909), Hubbard and Schwarz (1878), Downie and Arnett (1996); holotype from Isle Royale, MI

Habrocerus schwarzi Horn, 1877: Herman (2001), Hubbard and Schwarz (1878), Leng (1920), Downie and Arnett (1996), MSU-ARC

Subfamily Micropeplinae

Arrhenopeplus tesserula (Curtis, 1828): Herman (2001), Campbell (1984a), Hubbard and Schwarz (1878), Downie and Arnett (1996); most American sources (incl. BugGuide) do not use Arrhenopeplus and instead place this sp. in Micropeplus

Micropeplus browni Campbell, 1968: Herman (2001), Campbell (1984a), Downie and Arnett (1996)

Micropeplus laticollis Mäklin, 1853: Herman (2001), Campbell (1984a), Downie and Arnett (1996), MSU-ARC

Micropeplus sculptus LeConte, 1863: Herman (2001), Campbell (1984a), Downie and Arnett (1996)

Subfamily Mycetoporinae

Bobitobus cinctus (Gravenhorst, 1802): Herman (2001), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Bobitobus fungicola Campbell, 1982: Herman (2001), Campbell (1982a), Downie and Arnett (1996), MSU-ARC

Bobitobus kelleyi (Malkin, 1944): Herman (2001), Campbell (1982a), Downie and Arnett (1996), MSU-ARC

??Bobitobus obsoletus (Say, 1832): Hubbard and Schwarz (1878) as Bolitobius obsoletus Say

Bolitobius cingulatus Mannerheim, 1830: Wolcott (1909), Hubbard and Schwarz (1878), Ruesink and Parsons (2023b), MSU-ARC

Bryophacis smetanai Campbell, 1993: Herman (2001), Campbell (1993), Downie and Arnett (1996)

Bryoporus rufescens LeConte, 1863: Herman (2001), Leng (1920), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Bryoporus testaceus LeConte, 1863: Herman (2001), Campbell (1993), Hubbard and Schwarz (1878), Downie and Arnett (1996)

Carphacis effrenatus Herman, 2001: Herman (2001), Andrews (1923), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Carphacis nepigonensis (Bernhauer, 1912): Herman (2001), Downie and Arnett (1996), MSU-ARC

Carphasis dimidiatus (Erichson, 1839): Hubbard and Schwarz (1878), Ruesink and Parsons (2023b), MSU-ARC

Ischnosoma fimbriatum Campbell, 1991: Herman (2001), Campbell (1991), Downie and Arnett (1996); GBIF has this as a jr. syn. of I. longicorne Mäklin, BugGuide has fimbriatum as valid

Ischnosoma flavicolle (LeConte, 1863): Herman (2001), Schwarz (1876), Leng (1920), MSU-ARC

Ischnosoma pictum (Horn, 1877): Herman (2001), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Ischnosoma splendidum (Gravenhorst, 1806): Herman (2001), Downie and Arnett (1996), MSU-ARC

Ischnosoma virginicum (Bernhauer, 1917): Herman (2001), Campbell (1991), Downie and Arnett (1996), MSU-ARC

Lordithon anticus (Horn, 1877): Herman (2001), Campbell (1982a), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Lordithon appalachianus Campbell, 1982: Herman (2001), Campbell (1982a), Downie and Arnett (1996)

Lordithon exoletus (Erichson, 1839): Herman (2001), Townsend (1889), Leng (1920)

Lordithon facilis (Casey, 1885): Herman (2001), Campbell (1982a), Evans (2014), MSU-ARC

Lordithon niger (Gravenhorst, 1802): Herman (2001), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Lordithon thoracicus (Fabricius, 1777): Herman (2001), Andrews (1923), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Mycetoporus americanus Erichson, 1839: Herman (2001), Andrews (1923), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Mycetoporus consors LeConte, 1863: Herman (2001), Leng (1920), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Mycetoporus inquisitus Casey, 1885: Herman (2001), Campbell (1991), Downie and Arnett (1996), MSU-ARC

Mycetoporus lepidus (Gravenhorst, 1806): Herman (2001), Schwarz (1876), Hubbard and Schwarz (1878)

Mycetoporus lucidulus LeConte, 1863: Herman (2001), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Mycetoporus rufohumeralis Campbell, 1991: Herman (2001), Campbell (1991), Downie and Arnett (1996)

Mycetoporus rugosus Hatch, 1957: Herman (2001), Campbell (1991); GBIF has this as a jr. syn. of Mycetoporus montanus Luze, 1901

Subfamily Olisthaerinae

Olisthaerus megacephalus (Zetterstedt, 1828): Wolcott (1909), Hubbard and Schwarz (1878), LeConte (1850); Syn. laticeps described from St. Ignace by LeConte (1850)

Olisthaerus substriatus (Paykull, 1790): Wolcott (1909), Hubbard and Schwarz (1878), LeConte (1850); syn. nitidus described from Eagle Harbor by LeConte (1850)

Subfamily Osoriinae

Tribe Eleusinini

Eleusis pallida (LeConte, 1863): Herman (2001), Schwarz (1876), Hubbard and Schwarz (1878), MSU-ARC

Tribe Thoracophorini

Lispinodes undescribed species: Newton et al. (2001); recognized as a distinct species, but as of 2020 it remains undescribed as far as we know

Prolibia laevicauda (LeConte, 1866): Herman (2001), Downie and Arnett (1996)

Thoracophorus costalis (Erichson, 1840): Herman (2001), Hubbard and Schwarz (1878), MSU-ARC

Subfamily Oxyporinae

Oxyporus femoralis femoralis Gravenhorst, 1802: Herman (2001), Campbell (1969), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Oxyporus lateralis Gravenhorst, 1802: Herman (2001), Campbell (1969), Hanley (1993), Hubbard and Schwarz (1878), MSU-ARC; placed in genus Pseudoxyporus by GBIF

Oxyporus lepidus LeConte, 1877: Campbell (1969), Hanley (1993), Herman (2001), Downie and Arnett (1996)

Oxyporus major Gravenhorst, 1806: Hanley (1993), MSU-ARC

Oxyporus occipitalis Fauvel, 1864: Campbell (1969), Hanley (1993), Herman (2001), Downie and Arnett (1996), MSU-ARC; placed in genus Pseudoxyporus by GBIF

Oxyporus quinquemaculatus LeConte, 1863: Campbell (1969), Hanley (1993), Herman (2001), Andrews (1923), Downie and Arnett (1996), MSU-ARC; placed in genus Pseudoxyporus by GBIF

Oxyporus rufipennis LeConte, 1863: Campbell (1969), Hubbard and Schwarz (1878), Herman (2001), Andrews (1923), Downie and Arnett (1996), MSU-ARC

Oxyporus stygicus Say, 1831: Campbell (1969), Hubbard and Schwarz (1878), Herman (2001), Downie and Arnett (1996), MSU-ARC

Oxyporus vittatus vittatus Gravenhorst, 1802: Campbell (1969), Hanley (1993), Hubbard and Schwarz (1878), Herman (2001), MSU-ARC

Subfamily Phloeocharinae

Charhyphus picipennis (LeConte, 1863): Herman (2001), Hubbard and Schwarz (1878)

Subfamily Piestinae

Siagonium americanum (Melsheimer, 1844): Herman (2001), Hubbard and Schwarz (1878)

Siagonium punctatum (LeConte, 1866): Herman (2001)

Subfamily Proteininae

Tribe Proteinini

Megarthrus americanus Sachse, 1852: Cuccodoro and Löbl (1996), Andrews (1923)

Megarthrus excisus LeConte, 1863: Cuccodoro and Löbl (1996), Andrews (1923), Hubbard and Schwarz (1878)

Megarthrus pecki Cuccodoro and Löbl, 1996: Cuccodoro and Löbl (1996), Herman (2001)

Proteinus atomarius Erichson, 1840: Herman (2001), Ruesink and Parsons (2023b), MSU-ARC; a European sp.; Herman (2001) was citing Fauvel (1878) who considered parvulus a synonym, and Fauvel's info came from Schwarz's record for parvulus from Detroit, so Herman (2001) is not a valid report for atomarius.

Proteinus parvulus LeConte, 1863: Hubbard and Schwarz (1878); described from Lake Superior and known from across Canada, Hubbard and Schwarz (1878) report it from Detroit

Subfamily Pseudopsinae

Pseudopsis subulata Herman, 1975: Herman (2001), Downie and Arnett (1996)

Subfamily Xantholininae

Tribe Diochini

Diochus schaumi Kraatz, 1860: Herman (2001), Brunke et al. (2011), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Tribe Othiini

Atrecus americanus (Casey, 1906): Herman (2001), Smetana (1982), Downie and Arnett (1996), MSU-ARC

Tribe Xantholinini

Gyrohypnus campbelli Smetana, 1982: Herman (2001), Smetana (1982), Downie and Arnett (1996)

Gyrohypnus fracticornis (O.F. Müller, 1776): Herman (2001), Smetana (1982), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC

Hypnogyra gularis (LeConte, 1880): Herman (2001), Smetana (1982), Casey (1906), Hatch (1925), Downie and Arnett (1996)

Hypnogyra micans (Casey, 1906): Herman (2001), Smetana (1990)

Leptacinus batychrus (Gyllenhal, 1827): Herman (2001), Smetana (1982), Downie and Arnett (1996)

Leptacinus intermedius Donisthorpe, 1936: Ruesink and Parsons (2023b), MSU-ARC

Leptacinus pusillus (Stephens, 1833): Herman (2001), Smetana (1982), Downie and Arnett (1996), MSU-ARC

Lithocharodes longicollis (LeConte, 1863): Herman (2001), Smetana (1982), Hubbard and Schwarz (1878), Downie and Arnett (1996)

Neohypnus beckeri Smetana, 1982: Smetana (1982), Downie and Arnett (1996), MSU-ARC

Neohypnus emmesus (Gravenhorst, 1802): Herman (2001), Smetana (1982), Andrews (1916), Hubbard and Schwarz (1878), MSU-ARC

Neohypnus hamatus (Say, 1830): Herman (2001), Smetana (1982), Hatch (1925), Downie and Arnett (1996), MSU-ARC

Neohypnus lineatus (Wendeler, 1924): Herman (2001), Smetana (1982), MSU-ARC

Neohypnus melanops (Casey, 1906): Herman (2001), Smetana (1982)

Neohypnus monstrosus Smetana, 1982: Herman (2001), Smetana (1982), MSU-ARC

Neohypnus obscurus (Erichson, 1839): Herman (2001), Smetana (1982), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Neohypnus pusillus (Sasche, 1852): Herman (2001), Smetana (1982), Downie and Arnett (1996)

Nudobius cephalus (Say, 1830): Herman (2001), Smetana (1982), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC

Nudobius singularis Smetana, 1982: Herman (2001), Smetana (1982), Downie and Arnett (1996)

Phacophallus pallidipennis (Motschulsky, 1858): Ruesink and Parsons (2023b), MSU-ARC; slight chance the MSU specimens could be P. parumpunctatus instead

Stenistoderus rubripennis (LeConte, 1880): Herman (2001), Smetana (1982), Downie and Arnett (1996), MSU-ARC

Stictolinus flavipes (LeConte, 1863): Herman (2001), Smetana (1982), Downie and Arnett (1996), MSU-ARC

Xestolinus abdominalis Casey, 1906: Herman (2001), Smetana (1982), Downie and Arnett (1996), MSU-ARC

The following species were also reported from Michigan, but we consider them to be adventive, misidentifications or other errors:

Subfamily Mycetoporinae

Lordithon antennatus Campbell, 1982: KU:SEMC (Washtenaw Co.); a Mexican sp., should not be in MI

Lordithon trinotatus (Erichson, 1839): Hatch (1925), Hubbard and Schwarz (1878) as Bolitobius trinotatus Er.; a Eursian sp. that prob. does not occur in N.A.

Mycetoporus humidus (Say, 1832): Hatch (1925); humidus is a nomen dubium (Campbell 1991)

Subfamily Pseudopsinae

Pseudopsis columbica Fauvel, 1865: Leng (1920); a South American sp., cannot be in MI

Pseudopsis sulcata Newman, 1834: Hubbard and Schwarz (1878), Schwarz (1876); a European sp., does not occur in NA

Subfamily Xantholininae

Tribe Othiini

Atrecus pilicornis (Paykull, 1790): Hubbard and Schwarz (1878) as Baptolinus pilicornis Payk.; a European sp., does not occur in NA

This family list was last updated: April 28, 2023