Introduction. The name of each listed species is followed by up to five sources that report the species as occurring in Michigan. Sources in all upper-case letters refer to specific insect collections: MSU-ARC refers to material held by the Michigan State University Arthropod Research Collection (data provided by G. Parsons). All others refer to collection data extracted from the Symbiota Collections of Arthropods Network (SCAN) database online at https://scan-bugs.org/. See the Methods page for a list of abbreviations and associated collections.

Species names preceded by a single '?' are considered probable (for occurring in Michigan) but uncertain records. Those preceded by '??' are considered possible but unlikely to occur in Michigan. Species that are fully accepted as having valid Michigan records are unmarked. Species reported in the literature that we consider to be erroneous Michigan records are in a separate list following the others.

SCARABAEIDAE

6 subfamilies, 28 tribes, 62 genera, 168 species

Subfamily Aphodiinae

Tribe Aegialiini

Aegialia conferta Horn, 1871: Hubbard and Schwarz (1878), Ruesink and Parsons (2024), MSU-ARC

Aegialia criddlei Brown, 1931: Gordon and Cartwright (1988), MSU-ARC

Aegialia lacustris LeConte, 1850: Gordon and Cartwright (1988), Stebnicka (1977), Hubbard and Schwarz (1878), MSU-ARC

Aegialia opifex Horn, 1887: Downie and Arnett (1996), FMNH:FMNHINS (Berrien Co.), MSU-ARC

Caelius humeralis (Brown, 1931): Kriska and Young (2002), Gordon and Cartwright (1988)

Caelius rufescens (Horn, 1887): Gordon and Cartwright (1988), Stebnicka (1977), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Rhysothorax rufus (Fabricius, 1792): Gordon and Cartwright (1988), Stebnicka (1977), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Tribe Aphodiini

Acrossus rubripennis (Horn, 1870): Gordon and Skelley (2007), Bezanson and Floate (2019), Egan and Moon (2013), MSU-ARC

Agoliinus leopardus (Horn, 1870): Bezanson and Floate (2019), Rounds and Floate (2012), Hatch (1924), MSU-ARC

Agoliinus manitobensis (Brown, 1928): Bezanson and Floate (2019), Rounds and Floate (2012), MSU-ARC

Alloblackburneus lentus (Horn, 1870): Bezanson and Floate (2019), Wassmer (2020), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Alloblackburneus rubeolus (Palisot de Beauvois, 1805): Bezanson and Floate (2019), Gordon and Skelley (2007), Wassmer (2020), Dellacasa et al. (2011), MSU-ARC

??Aphodius fimetarius (Linnaeus, 1758): Bezanson and Floate (2019), Wassmer (2020), Andrews (1923), Hubbard and Schwarz (1878), Downie and Arnett (1996); "...what previously was widely reported in North America as A. fimetarius is now recognized as a cryptic species mix of A. fimetarius and Aphodius pedellus (De Geer) with overlapping distributions (Miraldo et al. 2014).

Aphodius pedellus (DeGeer, 1774): Miraldo et al. (2014), MSU-ARC

Blackburneus stercorosus (Melsheimer, 1845): Bezanson and Floate (2019), Wassmer (2020), Dellacasa et al. (2011), Hubbard and Schwarz (1878), MSU-ARC

Calamosternus granarius (Linnaeus, 1767): Bezanson and Floate (2019), Wassmer (2020), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC

Chilothorax distinctus (Müller, 1776): Bezanson and Floate (2019), Wassmer (2020), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC

Colobopterus erraticus (Linnaeus, 1758): Bezanson and Floate (2019), Rounds and Floate (2012), Wassmer (2020), Downie and Arnett (1996), MSU-ARC

Dialytellus tragicus (Schmidt, 1916): Gordon and Skelley (2007), Leng (1920), Hubbard and Schwarz (1878), Downie and Arnett (1996); GBIF has this a jr. syn. of D. humeralis (LeConte)

Dialytes striatulus (Say, 1825): Haack and Ruesink (2020), Hatch (1924), Hubbard and Schwarz (1878), MSU-ARC

Dialytes truncatus (Melsheimer, 1845): Bezanson and Floate (2019), Wassmer (2020), Downie and Arnett (1996), MSU-ARC

Dialytes ulkei (Horn, 1875): Ruesink and Parsons (2024), MSU-ARC

Diapterna hyperborea (LeConte, 1850): Hubbard and Schwarz (1878), Ruesink and Parsons (2024), MSU-ARC

Diapterna pinguis (Haldeman, 1848): Kriska and Young (2002), Hubbard and Schwarz (1878)

?Labarrus lividus (Olivier, 1789): BugGuide (Berrien Co.); "Taken from dog feces along Lake Michigan shore." (BugGuide)

Labarrus pseudolividus (Balthasar, 1941): Bezanson and Floate (2019), Wassmer (2020), MSU-ARC

Lechorodius terminalis (Say, 1823): Ruesink and Parsons (2024), MSU-ARC

Liothorax alternatus (Horn, 1870): Gordon (1977b), Dellacasa et al. (2007), Gordon and Skelley (2007), MSU-ARC

Melinopterus femoralis (Say, 1823): Gordon and Skelley (2007), Bezanson and Floate (2019), Kriska and Young (2002), MSU-ARC

Melinopterus prodromus (Brahm, 1790): Gordon and Skelley (2007), Bezanson and Floate (2019), Wassmer (2020), MSU-ARC

Oscarinus rusicola (Melsheimer, 1845): Bezanson and Floate (2019), Egan and Moon (2013), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC

Otophorus haemorrhoidalis (Linnaeus, 1758): Bezanson and Floate (2019), Wassmer (2020), MSU-ARC

Planolinellus vittatus (Say, 1825): Andrews (1923), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Pseudagolius bicolor (Say, 1823): Gordon and Skelley (2007), Bezanson and Floate (2019), Wassmer (2020), Hubbard and Schwarz (1878), MSU-ARC

Stenotothorax badipes (Melsheimer, 1845): Hubbard and Schwarz (1878), Ruesink and Parsons (2024), MSU-ARC; known also from IN

Teuchestes fossor (Linnaeus, 1758): Bezanson and Floate (2019), Wassmer (2014), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Tribe Eupariini

Ataenius abditus (Haldeman, 1848): Cartwright (1974), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Ataenius apicalis Hinton, 1936: Cartwright (1974)

Ataenius fattigi Cartwright, 1948: Cartwright (1974)

Ataenius gracilis (Melsheimer, 1845): Cartwright (1974), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Ataenius imbricatus (Melsheimer, 1845): Cartwright (1974), Bezanson and Floate (2019), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Ataenius punctifrons Cartwright, 1974: Cartwright (1974), Downie and Arnett (1996)

Ataenius spretulus (Haldeman, 1848): Cartwright (1974), Wassmer (2020), Downie and Arnett (1996), MSU-ARC

Ataenius strigatus (Say, 1823): Cartwright (1974), Wassmer (2014), Downie and Arnett (1996), MSU-ARC

Tribe Psammodiini

Platytomus atlanticus (Cartwright, 1948): Cartwright (1948), Downie and Arnett (1996); 6 paratypes from Detroit & Ganges, MI

Pleurophorus caesus (Creutzer, 1796): Ruesink and Parsons (2024), MSU-ARC

Rhyssemus germanus (Linnaeus, 1767): Gordon and Cartwright (1980), MSU-ARC

Subfamily Cetoniinae

Tribe Cetoniini

Euphoria fulgida (Fabricius, 1775): Orozco (2012), Andrews (1916), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Euphoria inda (Linnaeus, 1758): Orozco (2012), Andrews (1916), Hatch (1924), Hubbard and Schwarz (1878), MSU-ARC

Euphoria sepulcralis (Fabricius, 1801): Townsend (1889), iNaturalist Observations, Ruesink and Parsons (2024), MSU-ARC

Tribe Cremastocheilini

Cremastocheilus castaneae Knoch, 1801: Ruesink and Parsons (2024), MSU-ARC

?Cremastocheilus harrisii Kirby, 1827: Downie and Arnett (1996), Evans (2014); MI record needs confirmation

Cremastocheilus knochii LeConte, 1853: Ruesink and Parsons (2024), MSU-ARC

Cremastocheilus wheeleri LeConte, 1876: Ruesink and Parsons (2024), MSU-ARC

Tribe Gymnetini

Cotinis nitida (Linnaeus, 1764): Priest (2018), BugGuide, MSU-ARC

Tribe Osmodermini

Osmoderma eremicola (Knoch, 1801): Howden (1968), Hatch (1924), Downie and Arnett (1996), BugGuide, MSU-ARC

Osmoderma scabra (Palisot de Beauvois, 1805): Howden (1968), Hatch (1924), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC

Tribe Trichiini

Gnorimella maculosa (Knoch, 1801): Majka (2010), Howden (1968), Hubbard and Schwarz (1878), MSU-ARC

Trichiotinus affinis (Gory and Percheron, 1833): Kriska and Young (2002), Howden (1968), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Trichiotinus assimilis (Kirby, 1837): Howden (1968), BugGuide, MSU-ARC

Trichiotinus bibens (Fabricius, 1775): Kriska and Young (2002), Howden (1968), Downie and Arnett (1996), MSU-ARC

Trichiotinus piger (Fabricius, 1775): Howden (1968), Clampitt et al. (1996), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC

Trichiotinus viridans (Kirby, 1837): Howden (1968), BugGuide, MSU-ARC

Tribe Valgini

Valgus canaliculatus (Olivier, 1789): Jameson and Swoboda (2005), Townsend (1889), MSU-ARC

Valgus hemipterus (Linnaeus, 1758): Mahar and Oemke (1981), Jameson and Swoboda (2005), BugGuide, MSU-ARC

Valgus seticollis (Palisot de Beauvois, 1805): Townsend (1889), Evans (2014), Ruesink and Parsons (2024), MSU-ARC

Subfamily Dynastinae

Tribe Cyclocephalini

Cyclocephala borealis Arrow, 1911: Moore et al. (2018), Ratcliffe and Cave (2017)

Cyclocephala lurida lurida Bland, 1863: Moore et al. (2018), iNaturalist Observations, MSU-ARC

Dyscinetus morator (Fabricius, 1798): Moore et al. (2018), Townsend (1889), MSU-ARC

Tribe Dynastini

?Dynastes tityus (Linnaeus, 1763): Ratcliffe and Cave (2017); don't know where they got their record

Tribe Oryctini

Xyloryctes jamaicensis (Drury, 1773): Hardy (1991), Andrews (1916), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC; includes references to X. satyrus

Tribe Pentodontini

Aphonus densicauda Casey, 1915: Ratcliffe and Cave (2017)

Aphonus tridentatus (Say, 1823): Hardy (1991), Ratcliffe and Cave (2017), Andrews (1916), Leng (1920), MSU-ARC

Tomarus gibbosus (DeGeer, 1774): Hardy (1991), Ratcliffe and Cave (2017), Andrews (1916), MSU-ARC

Tomarus relictus (Say, 1825): Hardy (1991), Ratcliffe and Cave (2017), Andrews (1916), Hubbard and Schwarz (1878), MSU-ARC; GBIF places it in the genus Ligyrus

Tribe Phileurini

?Phileurus valgus (Olivier, 1789): Ratcliffe and Cave (2017); don't know where they got their record

Subfamily Melolonthinae

Tribe Dichelonychini

Dichelonyx albicollis Burmeister, 1855: Haack and Ruesink (2020), Andrews (1923), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Dichelonyx backii backii Kirby, 1837: Haack and Ruesink (2020), Downie and Arnett (1996), MSU-ARC

Dichelonyx diluta Fall, 1901: Kriska and Young (2002), Saylor (1945), Cornell (1972), Leng (1920), MSU-ARC

Dichelonyx elongatula (Schönherr, 1817): Haack and Ruesink (2020), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC; see comments under D. linearis

??Dichelonyx fuscula LeConte, 1856: Andrews (1923), Hubbard and Schwarz (1878); a more southern sp., MI record needs confirmation

Dichelonyx kirbyi Brown, 1946: Ruesink and Parsons (2024), MSU-ARC; MSU collection has 11 specimens, all from Dickinson Co., that we think are this species

Dichelonyx linearis (Gyllenhal, 1817): BugGuide, iNaturalist Observations, Ruesink and Parsons (2024), MSU-ARC; In his unpublished thesis Cornell (1972) considered this to be a synonym of D. elongatula as he could not find any consistent differences to separate the two. BugGuide says "Morphological differences between D. linearis and D. elongatulus are not well defined. Brown (1946) species concepts and geographic ranges are followed here where D. linearis occurs inland and D. elongatulus occurs in coastal areas (roughly the Maritimes through to New England)." MSU collection has specimens matching both nominal species.

Dichelonyx subvittata LeConte, 1856: Cornell (1972), Andrews (1923), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Tribe Diplotaxini

Diplotaxis atlantis Fall, 1909: Vaurie (1956), Vaurie (1960), Downie and Arnett (1996)

Diplotaxis harperi Blanchard, 1850: Vaurie (1960), MSU-ARC; many individuals of harperi are not separable externally from fulva.... (Vaurie, 1960)

??Diplotaxis liberta (Germar, 1824): Adams (1909), Hubbard and Schwarz (1878); an east coast & SE sp., MI records doubtful

Diplotaxis obscura LeConte, 1859: Vaurie (1960)

Diplotaxis sordida (Say, 1825): Vaurie (1956), Andrews (1923), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Diplotaxis tristis Kirby, 1837: Vaurie (1956), Andrews (1923), Downie and Arnett (1996), MSU-ARC

Tribe Hopliini

Hoplia modesta Haldeman, 1843: Hardy (1977), Andrews (1923), Downie and Arnett (1996), MSU-ARC

Hoplia trifasciata Say, 1825: Hardy (1977), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC

??Hoplia trivialis Harold, 1869: Downie and Arnett (1996), BugGuide; we suspect that most or all MI records for trivialis refer to trifasciata

Tribe Macrodactylini

??Macrodactylus angustatus(Palisot de Beauvois, 1805): OMNH:RINVRT (Cheboygan Co.); not likely to be that far north

Macrodactylus subspinosus (Fabricius, 1775): Clampitt et al. (1996), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC

Tribe Melolonthini

Amphimallon majale (Razoumowsky, 1789): Renkema et al. (2006), Smitley (1996), MSU-ARC

Phyllophaga anxia (LeConte, 1850): Luginbill and Painter (1953), Cantrall (1965), Andrews (1923), Downie and Arnett (1996), MSU-ARC

Phyllophaga balia (Say, 1825): Luginbill and Painter (1953), Cantrall (1965), MSU-ARC

?Phyllophaga barda (Horn, 1887): Downie and Arnett (1996); plausible, but where did they get the record?

Phyllophaga bipartita (Horn, 1887): Luginbill and Painter (1953), Downie and Arnett (1996)

Phyllophaga crenulata (Frölich, 1792): Luginbill and Painter (1953), Wellso and Fischer (1971), Cantrall (1965), Hubbard and Schwarz (1878), MSU-ARC

Phyllophaga drakii (Kirby, 1837): Luginbill and Painter (1953), Cantrall (1965), Andrews (1923), MSU-ARC

Phyllophaga fervida (Fabricius, 1775): Luginbill and Painter (1953), Kriska and Young (2002), Adams (1909), Downie and Arnett (1996), MSU-ARC

Phyllophaga forsteri (Burmeister, 1855): Luginbill and Painter (1953), Andrews (1916), Cantrall (1965), Downie and Arnett (1996), MSU-ARC

Phyllophaga fraterna Harris, 1841: Luginbill and Painter (1953), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Phyllophaga fusca (Frölich, 1792): Luginbill and Painter (1953), Cantrall (1965), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Phyllophaga futilis (LeConte, 1850): Luginbill and Painter (1953), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC

Phyllophaga gracilis (Burmeister, 1855): Wellso and Fischer (1971), Cantrall (1965), Andrews (1916), MSU-ARC

Phyllophaga hirticula (Knoch, 1801): Luginbill and Painter (1953), Cantrall (1965), Hubbard and Schwarz (1878), MSU-ARC

Phyllophaga hornii (Smith, 1889): Downie and Arnett (1996); plausible, but where did they get the record?

Phyllophaga ilicis (Knoch, 1801): Luginbill and Painter (1953), Cantrall (1965), Hubbard and Schwarz (1878), MSU-ARC

Phyllophaga implicita (Horn, 1887): Luginbill and Painter (1953), Smith (2012), MSU-ARC

Phyllophaga inversa (Horn, 1887): Luginbill and Painter (1953), MSU-ARC

Phyllophaga longispina (Smith, 1889): Luginbill and Painter (1953), Cantrall (1965), Leng (1920), Downie and Arnett (1996)

Phyllophaga marginalis (LeConte, 1856): Luginbill and Painter (1953), Cantrall (1965), Leng (1920), MSU-ARC

Phyllophaga micans (Knoch, 1801): Downie and Arnett (1996), MSU-ARC; ID of MSU specimen confirmed by GLP

Phyllophaga nitida (LeConte, 1856): Luginbill and Painter (1953), Cantrall (1965), MSU-ARC

Phyllophaga prunina (LeConte, 1856): Luginbill and Painter (1953), Cantrall (1965), Townsend (1893), Leng (1920), MSU-ARC

??Phyllophaga quercus (Knoch, 1801): Hubbard and Schwarz (1878) as Endrosa quercus Kn.; MI record needs confirmation, known from IN & OH.

Phyllophaga rubiginosa (LeConte, 1856): Luginbill and Painter (1953), Kriska and Young (2002), Downie and Arnett (1996)

Phyllophaga rugosa (Melsheimer, 1845): Luginbill and Painter (1953), Cantrall (1965), Wellso and Fischer (1971), Andrews (1923), MSU-ARC

Phyllophaga tristis (Fabricius, 1781): Luginbill and Painter (1953), Cantrall (1965), Townsend (1893), Hubbard and Schwarz (1878), MSU-ARC

Phyllophaga vilifrons (LeConte, 1856): Luginbill and Painter (1953), Cantrall (1965), Downie and Arnett (1996), MSU-ARC

Tribe Sericini

Maladera castanea (Arrow, 1913): Wassmer (2020), Smitley (1996), BugGuide

Serica atracapilla (Kirby, 1837): Dawson (1952), MSU-ARC

Serica blatchleyi Dawson, 1932: Ruesink and Parsons (2024), MSU-ARC

??Serica campestrisDawson, 1919: OMNH:RINVRT; Washtenaw Co.

Serica georgiana lecontei Dawson, 1921: Dawson (1952), AUMNH:ENT (Lake Co.), MSU-ARC

Serica intermixta Blatchley, 1910: Dawson (1952), Andrews (1916), MSU-ARC

Serica iricolor (Say, 1924): Wellso and Fischer (1971), Andrews (1916)

Serica loxia Dawson, 1920: Dawson (1952), Downie and Arnett (1996), MSU-ARC

Serica mystaca Dawson, 1922: Dawson (1952)

Serica parallela Casey, 1884: Dawson (1952), MSU-ARC

Serica sericea (Illiger, 1802): Dawson (1952), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC

Serica sponsa Dawson, 1919: Dawson (1952), Downie and Arnett (1996), MSU-ARC

Serica tristis LeConte, 1850: Dawson (1952), Andrews (1923), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Serica vespertina (Gyllenhal, 1817): Dawson (1952), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC

Subfamily Rutelinae

Tribe Anomalini

Anomala binotata (Gyllenhal, 1817): Hardy (1991), Townsend (1889), MSU-ARC

?Anomala flavipennis Burmeister, 1844: Downie and Arnett (1996), TTU:TTU-Z (Harrison, Clare Co.); known from IL, IN & OH, MI record needs confirmation

Anomala innuba (Fabricius, 1787): Wassmer (2020), Downie and Arnett (1996), MSU-ARC

Anomala lucicola (Fabricius, 1798): Hardy (1991), Downie and Arnett (1996), MSU-ARC

?Anomala ludoviciana Schaeffer, 1906: Andrews (1916), Downie and Arnett (1996); known from IL, IN & WI, MI record needs confirmation

??Anomala marginata (Fabricius, 1793): Downie and Arnett (1996); known from IN & OH, MI record needs confirmation

Anomala oblivia Horn, 1884: Hardy (1991), Downie and Arnett (1996), MSU-ARC

?Anomala orientalis (Waterhouse, 1875): BugGuide, iNaturalist Observations; placed in Exomala by some authors

Anomala undulata Melsheimer, 1845: Hardy (1991), Townsend (1893), Downie and Arnett (1996), MSU-ARC

Popillia japonica Newman, 1838: Clampitt et al. (1996), Haack and Ruesink (2020), MSU-ARC

Strigoderma arbicola (Fabricius, 1792): Bader (1992), Clampitt et al. (1996), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC

Tribe Rutelini

Cotalpa lanigera (Linnaeus, 1758): Ritcher (1966), Andrews (1923), Hubbard and Schwarz (1878), BugGuide, MSU-ARC

Pelidnota punctata (Linnaeus, 1758): Hardy (1991), Andrews (1916), Hubbard and Schwarz (1878), MSU-ARC

Subfamily Scarabaeinae

Tribe Ateuchini

Ateuchus histeroides Weber, 1801: Hubbard and Schwarz (1878), Robinson (1948), Downie and Arnett (1996), MSU-ARC

Tribe Coprini

Copris fricator (Fabricius, 1787): Bezanson and Floate (2019), Rounds and Floate (2012), Andrews (1916), Hubbard and Schwarz (1878), MSU-ARC

Copris minutus (Drury, 1773): Haack and Ruesink (2020), Hubbard and Schwarz (1878), MSU-ARC

Dichotomius carolinus (Linnaeus, 1767): Nielsen (1968)

Tribe Deltochilini

Canthon chalcites (Haldeman, 1843): Woodruff (1973)

Canthon pilularius (Linnaeus, 1758): Kriska and Young (2002), Andrews (1916), Downie and Arnett (1996), MSU-ARC

Canthon vigilans LeConte, 1858: Bezanson and Floate (2019), Leng (1920), Hubbard and Schwarz (1878), MSU-ARC

Melanocanthon nigricornis (Say, 1823): Bezanson and Floate (2019), Rounds and Floate (2012), Andrews (1916), Downie and Arnett (1996), MSU-ARC

Tribe Onthophagini

Onthophagus hecate hecate (Panzer, 1794): Bezanson and Floate (2019), Wassmer (2020), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC

Onthophagus nuchicornis (Linnaeus, 1758): Bezanson and Floate (2019), Rounds and Floate (2012), Howden and Cartwright (1963), Wassmer (2020), MSU-ARC

?Onthophagus orpheus pseudorpheus Howden & Cartwright, 1963: FMNH:FMNHINS; Berrien Co.

Onthophagus orpheus (Panzer, 1794): Bezanson and Floate (2019)

Onthophagus orpheuscanadensis (Fabricius, 1801): Downie and Arnett (1996), Howden and Cartwright (1963), MSU-ARC

Onthophagus pennsylvanicus Harold, 1871: Bezanson and Floate (2019), Howden and Cartwright (1963), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC

Onthophagus striatulus striatulus (Palisot de Beauvois, 1805): Hubbard and Schwarz (1878), Ruesink and Parsons (2024), MSU-ARC

Onthophagus taurus (Schreber, 1759): Bezanson and Floate (2019), Wassmer (2020), Parsons (2011)

Onthophagus tuberculifrons (Harold, 1871): Wassmer (2020), Howden and Cartwright (1963), Downie and Arnett (1996)

Tribe Phanaeini

Phanaeus vindex MacLeay, 1819: Nielsen (1968), Wassmer (2020), Townsend (1889), BugGuide, MSU-ARC; the Townsend (1889) record appears as P. carnifex L.

The following species were also reported from Michigan, but we consider them to be adventive, misidentifications or other errors:

Subfamily Aphodiinae

Tribe Aegialiini

Aegialia cylindrica (Eschscholtz, 1822): Hatch (1924); a west coast sp., cannot be in MI

Tribe Aphodiini

Aphodius fasciatus (Olivier, 1789): Hatch (1924); a European sp., does not occur in NA

Diapterna hamata (Say, 1824): Andrews (1916), Andrews (1923) as Aphodius hamatus Say; a western sp. that should not be in MI

Tribe Eupariini

Ataenius cognatus (LeConte, 1858): Andrews (1916); a SW species, cannot be in MI. Cartwright (1974): "Specimens from northern and eastern localities previously considered to be Ataenius cognatus are usually A. spretulus or A. strigatus."

Ataenius miamii Cartwright, 1934: Downie and Arnett (1996); a SE coastal sp., should not occur in MI

Ataenius stercorator (Fabricius, 1775): Hubbard and Schwarz (1878); a S. Amer. sp., does not occur in NA

Subfamily Cetoniinae

Tribe Trichiini

Trichiotinus lunulatus (Fabricius, 1775): Andrews (1916), Hubbard and Schwarz (1878) as Trichius viridulus Fabr.; a southern sp.

Subfamily Melolonthinae

Tribe Diplotaxini

Diplotaxis frondicola (Say, 1825): Hubbard and Schwarz (1878); known from southern parts of OH, IN & IL, should not be in MI

Tribe Melolonthini

Phyllophaga albina (Burmeister, 1855): Andrews (1916) as Lachnosterna albina Burm.; only two American records known

Phyllophaga hirsuta (Knoch, 1801): Hubbard and Schwarz (1878), TAMU:TAMUIC (Hillsdale Co.) as Lachnosterna hirsuta Kn.; an east coast & SE sp., should not be in MI

Tribe Sericini

Serica evidens Blatchley, 1919: Andrews (1916), Andrews (1923) as Serica carinata Blatchley; S. evidens is a SE species with northern limits just north of the Ohio River

Subfamily Scarabaeinae

Tribe Deltochilini

Canthon imitator Brown, 1946: FMNH:FMNHINS; Cheboygan; a southern plains sp.

This family list was last updated: March 10, 2023