Introduction. The name of each listed species is followed by up to five sources that report the species as occurring in Michigan. Sources in all upper-case letters refer to specific insect collections: MSU-ARC refers to material held by the Michigan State University Arthropod Research Collection (data provided by G. Parsons). All others refer to collection data extracted from the Symbiota Collections of Arthropods Network (SCAN) database online at https://scan-bugs.org/.  See the Methods page for a list of abbreviations and associated collections.

Species names preceded by a single '?' are considered probable (for occurring in Michigan) but uncertain records. Those preceded by '??' are considered possible but unlikely to occur in Michigan. Species that are fully accepted as having valid Michigan records are unmarked.  Species reported in the literature that we consider to be erroneous Michigan records are in a separate list following the others.

Bousquet (2012) did a marvelous job with the terrestrial Adephaga for all of North America north of Mexico, and although his assignments of species to subfamilies and tribes differs from other sources for some species, we follow his nomenclature and usage except in two cases: (1) in the Cicindelinae we follow Pearson et al. (2015), and (2) we follow the advice of Peter W. Messer and use Paratachys instead of Tachys in the Bembidiini.

CARABIDAE

13 subfamilies, 38 tribes, 98 genera, 528 species

Subfamily Brachininae

Tribe Brachinini

Brachinus adustipennis Erwin, 1969: Bousquet (2012), Downie and Arnett (1996)      

Brachinus alternans Dejean, 1825: Bousquet (2012), Downie and Arnett (1996), Andrews (1916), MSU-ARC      

Brachinus americanus (LeConte, 1844): Bousquet (2012), Downie and Arnett (1996), MSU-ARC      

Brachinus cordicollis Dejean, 1826: Bousquet (2012), Downie and Arnett (1996), Andrews (1916), MSU-ARC      

Brachinus cyanipennis Say, 1823: Bousquet (2012), Downie and Arnett (1996), Andrews (1916), MSU-ARC      

Brachinus cyanochroaticus Erwin, 1969: Bousquet (2012), Downie and Arnett (1996)      

Brachinus fumans (Fabricius, 1781): Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Brachinus janthinipennis (Dejean, 1831): Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

??Brachinus kansanus  LeConte, 1863: BYU:BYUC; a more southern sp., should not be in MI

Brachinus medius Harris, 1828: Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Brachinus microamericanus Erwin, 1969: Bousquet (2012), Downie and Arnett (1996)      

Brachinus ovipennis LeConte, 1863: Bousquet (2012), Downie and Arnett (1996)      

Brachinus patruelis LeConte, 1844: Bousquet (2012), Downie and Arnett (1996)      

??Brachinus phaeocerus  Chaudoir, 1868: BYU:BYUC; a plains sp., should not be in MI

Brachinus quadripennis Dejean, 1825: Bousquet (2012), Downie and Arnett (1996), Andrews (1916), Hubbard and Schwarz (1878), MSU-ARC      

Brachinus sublaevis Chaudoir, 1868: Bousquet (2012), Downie and Arnett (1996), MSU-ARC      

Brachinus tenuicollis LeConte, 1844: Bousquet (2012), Downie and Arnett (1996), Andrews (1916), MSU-ARC      

Brachinus texanus Chaudoir, 1868: Bousquet (2012), Downie and Arnett (1996)      

Subfamily Broscinae

Tribe Broscini

Miscodera arctica (Paykull, 1798): Bousquet (2012), Andrews (1923)      

Subfamily Carabinae

Tribe Carabini

Calosoma calidum (Fabricius, 1775): Bousquet (2012), Andrews (1916), Hatch (1924), Hubbard and Schwarz (1878), MSU-ARC      

Calosoma externum (Say, 1823): Bousquet (2012), MSU-ARC      

Calosoma frigidum Kirby, 1837: Bousquet (2012), Petrillo and Witter (2009), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

?Calosoma sayi Dejean, 1826: iNaturalist Observations; plausible but should be verified

Calosoma scrutator (Fabricius, 1775): Bousquet (2012), Nielsen (1968), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Calosoma wilcoxi LeConte, 1847: Bousquet (2012), Andrews (1916), Downie and Arnett (1996), MSU-ARC      

Carabus goryi Dejean, 1831: Bousquet (2012), Petrillo and Witter (2009), Haack et al. (2000), MSU-ARC      

Carabus maeander maeander Fischer von Waldheim, 1820: Bousquet (2012), Andrews (1929), Leng and Beutenmüller (1894a), Hubbard and Schwarz (1878), MSU-ARC      

Carabus nemoralis nemoralis Müller, 1764: Bousquet (2012), BugGuide, MSU-ARC      

Carabus serratus Say, 1823: Bousquet (2012), Andrews (1923), Hatch (1924), BugGuide, MSU-ARC      

Carabus sylvosus Say, 1823: Bousquet (2012), Andrews (1923), BugGuide, MSU-ARC      

Carabus vinctus (Weber, 1801): Bousquet (2012), Hubbard and Schwarz (1878)      

Tribe Cychrini

Scaphinotus bilobus (Say, 1823): Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Scaphinotus elevatus flammeus Haldeman, 1844: Ruesink and Parsons (2022), MSU-ARC      ID of specimen in MSU-ARC by Peter W. Messer

??Sphaeroderus canadensis canadensis Chaudoir, 1861: Bousquet (2012); MI record needs confirmation

Sphaeroderus schaumii Chaudoir, 1861: Bousquet (2012), Downie and Arnett (1996), BugGuide, MSU-ARC      

Sphaeroderus stenostomus lecontei Dejean, 1826: Bousquet (2012), Petrillo and Witter (2009), Dunn (1984), Hubbard and Schwarz (1878)      

Subfamily Cicindelinae

Tribe Cicindelini

Cicindela duodecimguttata Dejean, 1825: Bousquet (2012), Pearson et al. (2015), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Cicindela formosa generosa Dejean, 1831: Bousquet (2012), Pearson et al. (2015), Nielsen (1967), Hubbard and Schwarz (1878), MSU-ARC      

??Cicindela hirticollis hirticollis Say, 1817: Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878); Pearson et al. (2015) has this subsp. in OH & IN right at the MI border

Cicindela hirticollis rhodensis Calder, 1916: Bousquet (2012), Pearson et al. (2015), Dunn (1984), Andrews (1923), MSU-ARC; some of the MSU-ARC specimens are possibly hybrids or intergrades between this subspecies and the nominate one

Cicindela limbalis Klug, 1834: Bousquet (2012), Pearson et al. (2015), Downie and Arnett (1996), Andrews (1929), MSU-ARC      

Cicindela longilabris Say, 1824: Bousquet (2012), Pearson et al. (2015), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Cicindela patruela patruela Dejean, 1825: Bousquet (2012), Downie and Arnett (1996), Pearson et al. (2015), Hubbard and Schwarz (1878), MSU-ARC      

Cicindela purpurea purpurea Olivier, 1790: Bousquet (2012), Downie and Arnett (1996), Pearson et al. (2015), Hubbard and Schwarz (1878), MSU-ARC      

Cicindela repanda repanda Dejean, 1825: Bousquet (2012), Downie and Arnett (1996), Pearson et al. (2015), Hubbard and Schwarz (1878), MSU-ARC      

Cicindela scutellaris lecontei Haldeman, 1853: Bousquet (2012), Pearson et al. (2015), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Cicindela sexguttata Fabricius, 1775: Bousquet (2012), Pearson et al. (2015), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

??Cicindela tranquebarica kirbyi LeConte, 1867: BYU:BYUC, OSU:OSU       ID needs verification, this western subsp. reaches the west end of Lake Superior

Cicindela tranquebarica tranquebarica Herbst, 1806: Bousquet (2012), Downie and Arnett (1996), Pearson et al. (2015), Hubbard and Schwarz (1878), MSU-ARC      

Cicindelidia punctulata punctulata (Olivier, 1790): Bousquet (2012), Clampitt et al. (1996), Pearson et al. (2015), Downie and Arnett (1996), MSU-ARC      

Ellipsoptera lepida (Dejean, 1831): Bousquet (2012), Nielsen (1967), Pearson et al. (2015), Downie and Arnett (1996), MSU-ARC      

Ellipsoptera macra macra (LeConte, 1856): Bousquet (2012), Pearson et al. (2015), Downie and Arnett (1996), BugGuide, MSU-ARC      

Subfamily Elaphrinae

Tribe Elaphrini

Blethisa hudsonica Casey, 1924: Bousquet (2012), MSU-ARC; "This species has passed under the name B. multipunctata (Linnaeus, 1758) or B. multipunctata aurata Fischer von Waldheim, 1828 in the North American literature." Bousquet (2012)

??Blethisa julii LeConte, 1863: Bousquet (2012), Bousquet and Larochelle (1993); "The record from Michigan (Bousquet and Larochelle 1993: 86), based on a specimen labeled from Lake Superior in CMNH, needs confirmation." Bousquet (2012)

Blethisa quadricollis Haldeman, 1847: Bousquet (2012), Andrews (1923), Downie and Arnett (1996), Hubbard and Schwarz (1878)      

Elaphrus americanus americanus Dejean, 1831: Bousquet (2012), Downie and Arnett (1996), MSU-ARC      

Elaphrus californicus Mannerheim, 1843: Bousquet (2012), MSU-ARC      

Elaphrus cicatricosus LeConte, 1847: Bousquet (2012)      

Elaphrus clairvillei Kirby, 1837: Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Elaphrus fuliginosus Say, 1830: Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Elaphrus olivaceus LeConte, 1863: Bousquet (2012), Dunn (1984), Hubbard and Schwarz (1878), MSU-ARC      

Elaphrus ruscarius Say, 1830: Bousquet (2012), Hatch (1924), Hubbard and Schwarz (1878), MSU-ARC      

Subfamily Harpalinae

Tribe Abacetini

Oxycrepis velocipes (Casey, 1918): Ruesink and Parsons (2022), MSU-ARC; MSU-ARC specimen ID by Peter W. Messer

Tribe Atranini

Atranus pubescens (Dejean, 1828): Bousquet (2012), Hubbard and Schwarz (1878)      

Tribe Chlaeniini

Chlaenius aestivus Say, 1823: Bousquet (2012)      

Chlaenius brevilabris LeConte, 1847: Bousquet (2012), Hatch (1924), MSU-ARC      

Chlaenius cordicollis Kirby, 1837: Bousquet (2012), Dunn (1984), Downie and Arnett (1996), Hubbard and Schwarz (1878)      

Chlaenius emarginatus Say, 1823: Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Chlaenius erythropus Germar, 1824: Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878)      

Chlaenius impunctifrons Say, 1823: Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Chlaenius laticollis Say, 1823: Bousquet (2012), Townsend (1889)      

Chlaenius lithophilus Say, 1823: Bousquet (2012), Andrews (1923), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Chlaenius nemoralis Say, 1823: Bousquet (2012), Andrews (1923), Downie and Arnett (1996), MSU-ARC      

Chlaenius niger Randall, 1838: Bousquet (2012), Andrews (1923), Downie and Arnett (1996), Hubbard and Schwarz (1878)      

Chlaenius pennsylvanicus pennsylvanicus Say, 1823: Bousquet (2012), Andrews (1923), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Chlaenius platyderus Chaudoir, 1856: Bousquet (2012), Andrews (1923), Downie and Arnett (1996)      

Chlaenius prasinus Dejean, 1826: Bousquet (2012), Downie and Arnett (1996), MSU-ARC      

Chlaenius purpuricollis Randall, 1838: Bousquet (2012)      

Chlaenius pusillus Say, 1823: Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Chlaenius sericeus (Forster, 1771): Bousquet (2012), Downie and Arnett (1996), Andrews (1929), Hubbard and Schwarz (1878), MSU-ARC      

??Chlaenius solitarius Say, 1823: Bousquet (2012), Andrews (1916), Downie and Arnett (1996), Hatch (1924), UA:UAIC; Dunn (1984) says this record is an error, see long note in Bousquet (2012) regarding this matter

Chlaenius tomentosus (Say, 1823): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Chlaenius tricolor tricolor Dejean, 1826: Bousquet (2012), Hatch (1924), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Tribe Ctenodactylini

Leptotrachelus dorsalis (Fabricius, 1801): Bousquet (2012), Clampitt et al. (1996), MSU-ARC      

Tribe Cyclosomini

Tetragonoderus fasciatus (Haldeman, 1843): Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Tribe Galeritini

Galerita bicolor (Drury, 1773): Bousquet (2012), Andrews (1929), Hatch (1924), MSU-ARC      

Galerita janus (Fabricius, 1792): Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Tribe Harpalini

Acupalpus canadensis Casey, 1924: Bousquet (2012)      

Acupalpus carus (LeConte, 1863): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Acupalpus hydropicus (LeConte, 1863): Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Acupalpus indistinctus Dejean, 1831: Bousquet (2012)      

Acupalpus nanellus Casey, 1914: Bousquet (2012), TAMU:TAMUIC      

Acupalpus partiarius (Say, 1823): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Acupalpus pauperculus Dejean, 1829: Bousquet (2012), Hatch (1924), MSU-ARC      

Acupalpus pumilus Lindroth, 1968: Bousquet (2012), MSU-ARC      

Acupalpus testaceus Dejean, 1829: Bousquet (2012), MSU-ARC      

Agonoleptus conjunctus (Say, 1823): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Agonoleptus rotundicollis (Haldeman, 1843): Bousquet (2012), MSU-ARC      

Agonoleptus thoracicus (Casey, 1914): Bousquet (2012)      

Amphasia interstitialis (Say, 1823): Bousquet (2012), Andrews (1916), Hubbard and Schwarz (1878), MSU-ARC      

Amphasia sericea (Harris, 1828): Bousquet (2012), Andrews (1923), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Anisodactylus agricola (Say, 1823): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Anisodactylus caenus (Say, 1823): Bousquet (2012)      

Anisodactylus carbonarius (Say, 1823): Bousquet (2012), Hatch (1924), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Anisodactylus discoideus Dejean, 1831: Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Anisodactylus furvus LeConte, 1863: Bousquet (2012)      

Anisodactylus harrisii LeConte, 1863: Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Anisodactylus kirbyi Lindroth, 1953: Bousquet (2012), MSU-ARC      

Anisodactylus laetus Dejean, 1829: Bousquet (2012), MSU-ARC      

Anisodactylus melanopus (Haldeman, 1843): Bousquet (2012), MSU-ARC      

Anisodactylus merula (Germar, 1824): Bousquet (2012), Downie and Arnett (1996), MSU-ARC      

Anisodactylus nigerrimus (Dejean, 1831): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Anisodactylus nigrita Dejean, 1829: Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Anisodactylus ovularis (Casey, 1914): Bousquet (2012)      

Anisodactylus pseudagricola Noonan, 1996: Bousquet (2012)      

Anisodactylus rusticus (Say, 1823): Bousquet (2012), Andrews (1923), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Anisodactylus sanctaecrucis (Fabricius, 1798): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Anisodactylus verticalis (LeConte, 1847): Bousquet (2012), Andrews (1916), Hubbard and Schwarz (1878), MSU-ARC      

Bradycellus atrimedeus (Say, 1823): Bousquet (2012), Hubbard and Schwarz (1878)      

Bradycellus badipennis (Haldeman, 1843): Bousquet (2012), Hatch (1924), Hubbard and Schwarz (1878), MSU-ARC      

Bradycellus congener (LeConte, 1847): Bousquet (2012), MSU-ARC      

Bradycellus insulsus (Casey, 1914): Bousquet (2012), MSU-ARC      

Bradycellus kirbyi (Horn, 1883): Bousquet (2012), MSU-ARC      

Bradycellus lecontei Csiki, 1932: Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Bradycellus lugubris (LeConte, 1847): Bousquet (2012), Downie and Arnett (1996)      

Bradycellus neglectus (LeConte, 1847): Bousquet (2012), Downie and Arnett (1996), MSU-ARC      

Bradycellus nigriceps LeConte, 1869: Bousquet (2012), MSU-ARC      

Bradycellus nigrinus (Dejean, 1829): Bousquet (2012), Hatch (1924), Hubbard and Schwarz (1878), MSU-ARC      

Bradycellus rupestris (Say, 1823): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Bradycellus semipubescens Lindroth, 1968: Bousquet (2012)      

Cratacanthus dubius (Palisot de Beauvois, 1811): Bousquet (2012), Andrews (1916)      

Dicheirotrichus cognatus (Gyllenhal, 1827): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878)      

Discoderus parallelus (Haldeman, 1843): Bousquet (2012), MSU-ARC      

Euryderus grossus (Say, 1830): Bousquet (2012), Hatch (1924), MSU-ARC      

Geopinus incrassatus (Dejean, 1829): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Harpalus affinis (Schrank, 1781): Bousquet (2012), Andrews (1923), BugGuide, MSU-ARC      

Harpalus caliginosus (Fabricius, 1775): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Harpalus compar LeConte, 1847: Bousquet (2012), Hatch (1924), Hubbard and Schwarz (1878), MSU-ARC      

Harpalus erraticus Say, 1823: Bousquet (2012), Andrews (1923), Downie and Arnett (1996), MSU-ARC      

Harpalus erythropus Dejean, 1829: Bousquet (2012), Andrews (1916), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Harpalus faunus Say, 1823: Bousquet (2012), Hatch (1924), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Harpalus fulvilabris Mannerheim, 1853: Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878)      

Harpalus fuscipalpis Sturm, 1818: Bousquet (2012), Hubbard and Schwarz (1878), MSU-ARC      

Harpalus herbivagus Say, 1823: Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Harpalus indianus Csiki, 1932: Bousquet (2012), MSU-ARC      

Harpalus indigens Casey, 1924: Bousquet (2012), Downie and Arnett (1996), MSU-ARC      

Harpalus innocuus LeConte, 1863: Bousquet (2012), Andrews (1923), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Harpalus laevipes Zetterstedt, 1828: Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Harpalus laticeps LeConte, 1850: Bousquet (2012), Dunn (1984), Petrillo and Witter (2009), Hubbard and Schwarz (1878), MSU-ARC      

Harpalus lewisii LeConte, 1865: Bousquet (2012), Haack et al. (2000), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Harpalus megacephalus LeConte, 1847: Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Harpalus nigritarsis Sahlberg, 1827: Bousquet (2012), Downie and Arnett (1996)      

Harpalus opacipennis (Haldeman, 1843): Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Harpalus paratus Casey, 1924: Bousquet (2012), Downie and Arnett (1996)      

Harpalus pensylvanicus (DeGeer, 1774): Bousquet (2012), Haack et al. (2000), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Harpalus plenalis Casey, 1914: Bousquet (2012)      

Harpalus providens Casey, 1914: Bousquet (2012), MSU-ARC      

Harpalus reversus Casey, 1924: Bousquet (2012), Downie and Arnett (1996)      

Harpalus rubripes (Duftschmid, 1924): MSU-ARC; ID by G. Parsons, confirmed by Peter W. Messer

Harpalus solitaris Dejean, 1829: Bousquet (2012), Downie and Arnett (1996)      

Harpalus somnulentus Dejean, 1829: Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

??Harpalus spadiceus Dejean, 1829: Hubbard and Schwarz (1878); an east coast & Appalachian sp., but is recorded from Ohio. MI record needs confirmation.

Harpalus vagans LeConte, 1865: Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Notiobia nitidipennis (LeConte, 1847): Bousquet (2012), Downie and Arnett (1996), MSU-ARC      

Notiobia sayi (Blatchley, 1910): Bousquet (2012), Andrews (1923), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Notiobia terminata (Say, 1823): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Ophonus puncticeps Stephens, 1828: Bousquet (2012), Clampitt et al. (1996), Purrington et al. (2000), MSU-ARC      

Philodes rectangulus (Chaudoir, 1868): Bousquet (2012)      

Selenophorus ellipticus Dejean, 1829: Bousquet (2012), MSU-ARC      

Selenophorus gagatinus Dejean, 1829: Bousquet (2012), MSU-ARC      

Selenophorus granarius Dejean, 1829: Messer and Raber (2021)      

Selenophorus hylacis (Say, 1823): Bousquet (2012), Andrews (1916), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Selenophorus opalinus (LeConte, 1863): Bousquet (2012), Andrews (1916), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Selenophorus planipennis LeConte, 1847: Bousquet (2012)      

Stenolophus carbo Bousquet, 1993: Bousquet (2012), Leng (1920), Downie and Arnett (1996)      

Stenolophus comma (Fabricius, 1775): Bousquet (2012), Dunn (1984), Hubbard and Schwarz (1878), MSU-ARC      

Stenolophus dissimilis Dejean, 1829: Bousquet (2012), Andrews (1916)      

Stenolophus fuliginosus Dejean, 1829: Bousquet (2012), Andrews (1916), Hubbard and Schwarz (1878), MSU-ARC      

Stenolophus fuscatus Dejean, 1829: Bousquet (2012), Hatch (1924)      

Stenolophus humidus Hamilton, 1893: Bousquet (2012)      

Stenolophus lecontei (Chaudoir, 1868): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Stenolophus lineola (Fabricius, 1775): Bousquet (2012), Hatch (1924), Hubbard and Schwarz (1878), MSU-ARC      

Stenolophus ochropezus (Say, 1823): Bousquet (2012), Dunn (1984), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Stenolophus plebejus Dejean, 1829: Bousquet (2012), Hatch (1924), Hubbard and Schwarz (1878)      

Trichotichnus autumnalis (Say, 1823): Bousquet (2012), Andrews (1916), Hubbard and Schwarz (1878), MSU-ARC      

Trichotichnus dichrous (Dejean, 1829): Bousquet (2012), Hubbard and Schwarz (1878), MSU-ARC      

Trichotichnus vulpeculus (Say, 1823): Bousquet (2012), MSU-ARC      

Xestonotus lugubris (Dejean, 1829): Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Tribe Helluonini

Helluomorphoides nigripennis (Dejean, 1831): Bousquet (2012)      

Helluomorphoides praeustus bicolor (Harris, 1828): Bousquet (2012), Townsend (1889), Downie and Arnett (1996), BugGuide, MSU-ARC      

Tribe Lebiini

Apenes lucidula lucidula (Dejean, 1831): Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878)      

Apenes sinuata (Say, 1823): Bousquet (2012), MSU-ARC      

Apristus subsulcatus (Dejean, 1826): Bousquet (2012)      

Axinopalpus biplagiatus (Dejean, 1825): Bousquet (2012), Hubbard and Schwarz (1878)      

Calleida punctata LeConte, 1846: Bousquet (2012), Clampitt et al. (1996), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Calleida purpurea (Say, 1823): Bousquet (2012), Andrews (1916), Hubbard and Schwarz (1878), MSU-ARC      

Coptodera aerata Dejean, 1825: Bousquet (2012)      

Cymindis americana Dejean, 1826: Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Cymindis borealis LeConte, 1863: Bousquet (2012)      

Cymindis cribricollis Dejean, 1831: Bousquet (2012), Andrews (1923), BugGuide, Hubbard and Schwarz (1878), MSU-ARC      

Cymindis limbata Dejean, 1831: Bousquet (2012), Hunting (2013), Hubbard and Schwarz (1878), MSU-ARC      

Cymindis neglecta Haldeman, 1843: Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Cymindis pilosa Say, 1823: Bousquet (2012), Hatch (1924), Hubbard and Schwarz (1878), MSU-ARC      

Cymindis platicollis platicollis (Say, 1823): Bousquet (2012), Hunting (2013), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Dromius piceus Dejean, 1831: Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Lebia analis Dejean, 1825: Bousquet (2012), Downie and Arnett (1996), MSU-ARC      

Lebia atriventris Say, 1823: Bousquet (2012), Andrews (1923), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Lebia bivittata (Fabricius, 1798): Bousquet (2012), Downie and Arnett (1996), MSU-ARC      

?Lebia divisa LeConte, 1850: Hubbard and Schwarz (1878); record needs confirmation, although Hubbard and Schwarz (1878) say theirs was taken by LeConte at Eagle Harbor, known from western IN & WI, but MI not listed in Bousquet (2012)

Lebia fuscata Dejean, 1825: Bousquet (2012), Andrews (1916), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Lebia grandis Hentz, 1830: Bousquet (2012), Andrews (1916), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Lebia marginicollis Dejean, 1825: Bousquet (2012), Downie and Arnett (1996), MSU-ARC      

Lebia moesta LeConte, 1850: Bousquet (2012), Hatch (1924), Hubbard and Schwarz (1878), MSU-ARC      

Lebia ornata Say, 1823: Bousquet (2012), Clampitt et al. (1996), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Lebia pectita Horn, 1885: Bousquet (2012), Evans (2014), Downie and Arnett (1996)      

Lebia pleuritica LeConte, 1846: Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Lebia pulchella Dejean, 1826: Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Lebia pumila Dejean, 1831: Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Lebia solea Hentz, 1830: Bousquet (2012), Andrews (1923), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Lebia tricolor tricolor Say, 1823: Bousquet (2012), Andrews (1923), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Lebia viridipennis Dejean, 1826: Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Lebia viridis Say, 1823: Bousquet (2012), Clampitt et al. (1996), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Lebia vittata (Fabricius, 1777): Bousquet (2012), Hatch (1924), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Microlestes linearis (LeConte, 1851): Bousquet (2012), Hatch (1924), Hubbard and Schwarz (1878), MSU-ARC      

Plochionus timidus Haldeman, 1843: Bousquet (2012), Hubbard and Schwarz (1878), MSU-ARC      

Syntomus americanus (Dejean, 1831): Bousquet (2012), Hubbard and Schwarz (1878), MSU-ARC      

Tribe Licinini

Badister flavipes flavipes LeConte, 1853: Bousquet (2012), Downie and Arnett (1996), MSU-ARC      

Badister grandiceps Casey, 1920: Bousquet (2012), Downie and Arnett (1996)      

Badister micans LeConte, 1844: Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Badister neopulchellus Lindroth, 1954: Bousquet (2012), BugGuide, MSU-ARC      

Badister notatus Haldeman, 1843: Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878)      

Badister obtusus LeConte, 1878: Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Badister parviceps Ball, 1959: Bousquet (2012)      

Badister pulchellus LeConte, 1847: Bousquet (2012), Andrews (1923), Hatch (1924), Hubbard and Schwarz (1878)      

Badister reflexus LeConte, 1880: Bousquet (2012), Andrews (1923), Leng (1920), MSU-ARC      

Badister transversus Casey, 1920: Bousquet (2012)      

Dicaelus elongatus Bonelli, 1813: Bousquet (2012), Downie and Arnett (1996), MSU-ARC      

Dicaelus politus Dejean, 1826: Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Dicaelus purpuratus purpuratus Bonelli, 1813: Bousquet (2012), Andrews (1916), Hubbard and Schwarz (1878), MSU-ARC      

Dicaelus sculptilis intricatus LeConte, 1873: Bousquet (2012), Downie and Arnett (1996)      

Dicaelus teter Bonelli, 1813: Bousquet (2012), Petrillo and Witter (2009), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Diplocheila assimilis (LeConte, 1844): Bousquet (2012), Downie and Arnett (1996), MSU-ARC      

Diplocheila impressicollis (Dejean, 1831): Bousquet (2012), Andrews (1923), BugGuide, Hubbard and Schwarz (1878), MSU-ARC      

Diplocheila major major (LeConte, 1847): Bousquet (2012), Andrews (1916), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Diplocheila obtusa (LeConte, 1847): Bousquet (2012), Andrews (1916), Downie and Arnett (1996), MSU-ARC      

Diplocheila striatopunctata (LeConte, 1844): Bousquet (2012), Andrews (1923), Downie and Arnett (1996), MSU-ARC      

Tribe Odacanthini

Colliuris pensylvanica (Linnaeus, 1758): Bousquet (2012), Andrews (1923), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Tribe Oodini

Lachnocrepis parallela (Say, 1830): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Oodes amaroides Dejean, 1831: Bousquet (2012), Andrews (1916), MSU-ARC      

Oodes brevis Lindroth, 1957: Bousquet (2012)      

Oodes fluvialis LeConte, 1863: Bousquet (2012), Andrews (1916), Hubbard and Schwarz (1878), MSU-ARC      

Stenocrepis cuprea (Chaudoir, 1843): Bousquet (2012)      

Tribe Panagaeini

Panagaeus fasciatus Say, 1823: Bousquet (2012)      

Tribe Pentagonicini

Pentagonica picticornis Bates, 1883: Bousquet (2012), MSU-ARC      

Tribe Perogonini

Perigona nigriceps (Dejean, 1831): Bousquet (2012), Hubbard and Schwarz (1878)      

Perigona pallipennis (LeConte, 1853): Bousquet (2012), Downie and Arnett (1996)      

Tribe Platynini

Agonum aeruginosum Dejean, 1828: Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Agonum affine Kirby, 1837: Bousquet (2012), Andrews (1923), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Agonum anchomenoides Randall, 1838: Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Agonum canadense Goulet, 1969: Bousquet (2012), MSU-ARC      

Agonum collare (Say, 1830): Bousquet (2012)      

??Agonum corvus (LeConte, 1860): Bousquet (2012); "Needs confirmation" Bousquet (2012)

Agonum crenistriatum (LeConte, 1863): Bousquet (2012), Andrews (1923), Hatch (1924), Hubbard and Schwarz (1878), MSU-ARC      

Agonum cupreum Dejean, 1831: Bousquet (2012), Andrews (1923), Hatch (1924), MSU-ARC      

Agonum cupripenne (Say, 1823): Bousquet (2012), Petrillo and Witter (2009), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Agonum decorum (Say, 1823): Bousquet (2012), Clampitt et al. (1996), Hubbard and Schwarz (1878), MSU-ARC      

Agonum errans (Say, 1823): Bousquet (2012), Dunn (1984), Hubbard and Schwarz (1878), MSU-ARC      

Agonum excavatum Dejean, 1828: Bousquet (2012), Andrews (1916), Hubbard and Schwarz (1878), MSU-ARC      

Agonum extensicolle (Say, 1823): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Agonum ferreum Haldeman, 1843: Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878)      

Agonum fidele Casey, 1920: Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878)      

Agonum galvestonicum (Casey, 1920): Bousquet (2012), MSU-ARC      

Agonum gratiosum (Mannerheim, 1853): Bousquet (2012), Dunn (1984), Hubbard and Schwarz (1878), MSU-ARC      

Agonum harrisii LeConte, 1846: Bousquet (2012), MSU-ARC      

Agonum lutulentum (LeConte, 1854): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Agonum melanarium Dejean, 1828: Bousquet (2012), Andrews (1923), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Agonum metallescens (LeConte, 1854): Bousquet (2012), Hubbard and Schwarz (1878)      

Agonum moerens Dejean, 1828: Bousquet (2012), Andrews (1923), MSU-ARC      

Agonum muelleri (Herbst, 1784): Bousquet (2012)      

Agonum mutatum (Gemminger & Harold, 1868): Bousquet (2012), Andrews (1923), Hatch (1924), Hubbard and Schwarz (1878), MSU-ARC      

Agonum nigriceps LeConte, 1846: Bousquet (2012), Hubbard and Schwarz (1878)      

Agonum nutans (Say, 1823): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Agonum octopunctatum (Fabricius, 1798): Bousquet (2012), BugGuide, Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Agonum palustre Goulet, 1969: Bousquet (2012), MSU-ARC      

Agonum piceolum (LeConte, 1879): Bousquet (2012)      

Agonum picicornoides Lindroth, 1966: Bousquet (2012), Downie and Arnett (1996)      

Agonum placidum (Say, 1823): Bousquet (2012), Dunn (1984), Hatch (1924), Hubbard and Schwarz (1878), MSU-ARC      

Agonum propinquum (Gemminger & Harold, 1868): Bousquet (2012), MSU-ARC      

Agonum punctiforme (Say, 1823): Bousquet (2012), Andrews (1916), MSU-ARC      

??Agonum quadrimaculatum (Horn, 1885): Bousquet (2012); "Needs confirmation" Bousquet (2012)

Agonum retractum LeConte, 1846: Bousquet (2012), Andrews (1923), MSU-ARC      

Agonum rufipes Dejean, 1828: Bousquet (2012), Andrews (1916), Andrews (1923)      

Agonum sordens Kirby, 1837: Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Agonum striatopunctatum Dejean, 1828: Bousquet (2012), MSU-ARC      

Agonum superioris Lindroth, 1966: Bousquet (2012)      

Agonum tenue (LeConte, 1854): Bousquet (2012), Dunn (1984), Andrews (1923), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Agonum thoreyi Dejean, 1828: Bousquet (2012), Clampitt et al. (1996), Dunn (1984), Hubbard and Schwarz (1878), MSU-ARC      

Agonum trigeminum Lindroth, 1954: Bousquet (2012), MSU-ARC      

Olisthopus micans LeConte, 1846: Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Olisthopus parmatus (Say, 1823): Bousquet (2012), Andrews (1916), Hubbard and Schwarz (1878)      

Oxypselaphus pusillus (LeConte, 1854): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Platynus cincticollis (Say, 1823): Bousquet (2012), Andrews (1916), MSU-ARC      

Platynus decentis (Say, 1823): Bousquet (2012), Petrillo and Witter (2009), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Platynus hypolithos (Say, 1823): Bousquet (2012), Clampitt et al. (1996), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Platynus mannerheimii (Dejean, 1828): Bousquet (2012)      

Platynus opaculus LeConte, 1863: Bousquet (2012), Andrews (1916), Downie and Arnett (1996), MSU-ARC      

Platynus tenuicollis (LeConte, 1846): Bousquet (2012), Andrews (1923), Hatch (1924), MSU-ARC      

Sericoda obsoleta (Say, 1823): Bousquet (2012), Hubbard and Schwarz (1878), MSU-ARC      

Sericoda quadripunctata (DeGeer, 1774): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Tribe Pterostichini

Cyclotrachelus freitagi Bousquet, 1993: Bousquet (2012), Downie and Arnett (1996)      

Cyclotrachelus seximpressus (LeConte, 1846): Bousquet (2012), Downie and Arnett (1996)      

Cyclotrachelus sodalis sodalis (LeConte, 1846): Bousquet (2012), Downie and Arnett (1996), MSU-ARC      

Gastrellarius honestus (Say, 1823): Bousquet (2012), Andrews (1923), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Lophoglossus scrutator (LeConte, 1846): Bousquet (2012), Andrews (1916), Hubbard and Schwarz (1878), MSU-ARC      

Myas coracinus (Say, 1823): Bousquet (2012), Dunn (1984), Townsend (1889), MSU-ARC      

Myas cyanescens Dejean, 1828: Bousquet (2012), Petrillo and Witter (2009), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Poecilus chalcites (Say, 1823): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Poecilus lucublandus (Say, 1823): Bousquet (2012), Petrillo and Witter (2009), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Pterostichus adoxus (Say, 1823): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Pterostichus adstrictus Eschscholtz, 1823: Bousquet (2012), Petrillo and Witter (2009), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Pterostichus brevicornis brevicornis (Kirby, 1837): Bousquet (2012)      

Pterostichus castor Goulet & Bousquet, 1983: Bousquet (2012)      

Pterostichus caudicalis (Say, 1823): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Pterostichus commutabilis (Motschulsky, 1866): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Pterostichus coracinus (Newman, 1838): Bousquet (2012), Dunn (1984), Petrillo and Witter (2009), Hubbard and Schwarz (1878), MSU-ARC      

Pterostichus corvinus (Dejean, 1828): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Pterostichus ebeninus (Dejean, 1828): Bousquet (2012)      

Pterostichus fallax (Dejean, 1828): Bousquet (2012)      

Pterostichus femoralis (Kirby, 1837): Bousquet (2012), Adams (1909), Hubbard and Schwarz (1878), MSU-ARC      

Pterostichus luctuosus (Dejean, 1828): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Pterostichus melanarius melanarius (Illiger, 1798): Bousquet (2012), Dunn (1984), Petrillo and Witter (2009), MSU-ARC      

Pterostichus mutus (Say, 1823): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Pterostichus novus Straneo, 1944: Bousquet (2012), Downie and Arnett (1996), MSU-ARC      

Pterostichus patruelis (Dejean, 1831): Bousquet (2012), Hubbard and Schwarz (1878), MSU-ARC      

Pterostichus pensylvanicus LeConte, 1873: Bousquet (2012), Haack et al. (2000), Petrillo and Witter (2009), Andrews (1916), MSU-ARC      

Pterostichus permundus (Say, 1830): Bousquet (2012), Andrews (1923), MSU-ARC      

Pterostichus praetermissus (Chaudoir, 1868): Bousquet (2012)      

Pterostichus punctatissimus (Randall, 1838): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878)      

Pterostichus relictus (Newman, 1838): Bousquet (2012), Wickham (1895), Andrews (1923), MSU-ARC      

Pterostichus stygicus (Say, 1823): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Pterostichus tenuis (Casey, 1924): Bousquet (2012), Downie and Arnett (1996)      

Pterostichus trinarius (Casey, 1918): Bousquet (2012)      

Pterostichus tristis (Dejean, 1828): Bousquet (2012), Dunn (1984), Petrillo and Witter (2009)      

??Stereocerus  haematopus  (Dejean, 1831): OSU:OSU; a far northern sp., unlikely to be in MI

Tribe Sphodrini

Calathus advena (LeConte, 1846): Bousquet (2012), Adams (1909), Hubbard and Schwarz (1878), MSU-ARC      

Calathus gregarius (Say, 1823): Bousquet (2012), Petrillo and Witter (2009), Andrews (1923), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Calathus ingratus Dejean, 1828: Bousquet (2012), Dunn (1984), Clampitt et al. (1996), MSU-ARC      

Calathus opaculus LeConte, 1854: Bousquet (2012), Andrews (1923), Downie and Arnett (1996), MSU-ARC      

Pseudamara arenaria (LeConte, 1847): Bousquet (2012), Andrews (1923), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Synuchus impunctatus (Say, 1823): Bousquet (2012), Petrillo and Witter (2009), Dunn (1984), Hubbard and Schwarz (1878), MSU-ARC      

Tribe Zabrini

Amara aenea (DeGeer, 1774): Bousquet (2012), BugGuide, MSU-ARC      

Amara aeneopolita Casey, 1918: Bousquet (2012), Downie and Arnett (1996)      

Amara angustata (Say, 1823): Bousquet (2012), Andrews (1923), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Amara angustatoides Hieke, 2000: Bousquet (2012)      

Amara apricaria (Paykull, 1790): Bousquet (2012), Andrews (1923), MSU-ARC      

Amara avida (Say, 1823): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

??Amara basillaris (Say, 1823): Andrews (1923); MI record needs confirmation

?Amara carinata (LeConte, 1847): U Alberta, UASM:UASM; should get this ID confirmed

Amara chalcea Dejean, 1828: Bousquet (2012), Andrews (1923)      

??Amara confusa LeConte, 1847: Andrews (1923); MI record needs confirmation

Amara convexa LeConte, 1847: Bousquet (2012), Andrews (1923), BugGuide, Hubbard and Schwarz (1878), MSU-ARC      

Amara crassispina LeConte, 1855: Bousquet (2012), Andrews (1923), MSU-ARC      

Amara cupreolata Putzeys, 1866: Bousquet (2012), Andrews (1923), MSU-ARC      

Amara ellipsis (Casey, 1918): Bousquet (2012), Haack and Ruesink (2020)      

??Amara erratica (Duftschmid, 1812): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), UA:UAIC; "Several state records (e.g., CA, IA, MI, MN, ND, NM, OR) listed in Bousquet and Larochelle (1993: 196) and taken from earlier authors probably refer to other species or need confirmation." Bousquet (2012)

Amara exarata Dejean, 1828: Bousquet (2012), Andrews (1916), Andrews (1923)      

Amara familiaris (Duftschmid, 1812): Bousquet (2012), MSU-ARC      

Amara flebilis (Casey, 1918): Bousquet (2012)      

Amara gibba (LeConte, 1847): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Amara hyperborea Dejean, 1831: Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878)      

Amara impuncticollis (Say, 1823): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

??Amara interstitialis Dejean, 1828: CSU:CSUC, Andrews (1923), Hubbard and Schwarz (1878); should get this ID confirmed, a far northern sp. that should not be this far south

Amara lacustris LeConte, 1855: Bousquet (2012)      

Amara laevipennis Kirby, 1837: Bousquet (2012), Downie and Arnett (1996)      

Amara latior (Kirby, 1837): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Amara littoralis Dejean, 1828: Bousquet (2012), Andrews (1923), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Amara lunicollis Schiodte, 1837: Bousquet (2012), Downie and Arnett (1996)      

Amara musculis (Say, 1823): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Amara neoscotica Casey, 1924: Bousquet (2012)      

Amara obesa (Say, 1823): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Amara otiosa Casey, 1918: Bousquet (2012)      

Amara pallipes Kirby, 1837: Bousquet (2012), Downie and Arnett (1996), MSU-ARC      

Amara patruelis Dejean, 1831: Bousquet (2012), MSU-ARC      

Amara pennsylvanica Hayward, 1908: Bousquet (2012), MSU-ARC      

Amara quenseli quenseli (Schönherr, 1806): Bousquet (2012), Andrews (1923), Downie and Arnett (1996), MSU-ARC      

Amara rubrica Haldeman, 1843: Bousquet (2012), Andrews (1923), BugGuide, MSU-ARC      

Amara schwarzi Hayward, 1908: Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878)      

Amara sinuosa (Casey, 1918): Bousquet (2012), Andrews (1923), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Amara tenax Casey, 1918: Bousquet (2012)      

??Amara torrida (Panzer, 1796): Andrews (1923); a circumpolar species, MI record needs confirmation

Amara turbata Casey, 1918: Bousquet (2012)      

Tribe Zuphiini

Zuphioides americanum (Dejean, 1831): Bousquet (2012), Downie and Arnett (1996)      

Subfamily Loricerinae

Tribe Loricerini

Loricera pilicornis pilicornis (Fabricius, 1775): Bousquet (2012), Andrews (1923), Dunn (1984), Hubbard and Schwarz (1878), MSU-ARC      

Subfamily Nebriinae

Tribe Nebriini

Nebria lacustris Casey, 1913: Bousquet (2012)      

Nebria pallipes Say, 1823: Bousquet (2012), Hubbard and Schwarz (1878)      

Nebria rufescens  castanipes (Kirby, 1837): Bousquet (2012)      

??Nebria suturalis LeConte, 1850: Andrews (1923); only MI record, needs confirmation. Bousquet (2012) has it across Canada.

Tribe Notiophilini

Notiophilus aeneus (Herbst, 1806): Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Notiophilus aquaticus (Linnaeus, 1758): Bousquet (2012), Hubbard and Schwarz (1878), MSU-ARC      

Notiophilus novemstriatus LeConte, 1847: Bousquet (2012), Downie and Arnett (1996)      

Notiophilus semistriatus Say, 1823: Bousquet (2012), Andrews (1923), Hatch (1924), Hubbard and Schwarz (1878), MSU-ARC      

Subfamily Omophroninae

Tribe Omophronini

Omophron americanum Dejean, 1831: Bousquet (2012), Dunn (1984), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Omophron nitidum LeConte, 1847: Ruesink and Parsons (2022), Andrews (1916), MSU-ARC; "The record from ... Charity Island in Michigan (Andrews 1916: 72) [is] probably based on misidentified O. americanum." Bousquet (2012)

Omophron robustum Horn, 1870: Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Omophron tessellatum Say, 1823: Bousquet (2012), Andrews (1923), BugGuide, Hubbard and Schwarz (1878), MSU-ARC      

Subfamily Patrobinae

Tribe Patrobini

Patrobus cinctus Motschulsky, 1860: Bousquet (2012)      

Patrobus foveocollis (Eschscholtz, 1823): Bousquet (2012), Hubbard and Schwarz (1878), Leng (1920), MSU-ARC      

Patrobus longicornis (Say, 1823): Bousquet (2012), Andrews (1923), BugGuide, Hubbard and Schwarz (1878), MSU-ARC      

Patrobus septentrionis septentrionis Dejean, 1828: Bousquet (2012), Downie and Arnett (1996)      

Patrobus stygicus Chaudoir, 1872: Bousquet (2012)      

Subfamily Psydrinae

Tribe Psydrini

Nomius pygmaeus (Dejean, 1831): Bousquet (2012), Andrews (1923), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Psydrus piceus LeConte, 1846: Bousquet (2012), Hubbard and Schwarz (1878)      

Subfamily Scaritinae

Tribe Clivinini

Aspidoglossa subangulata (Chaudoir, 1843): Bousquet (2012)      

Clivina americana Dejean, 1831: Bousquet (2012), Leng (1920), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Clivina dentipes Dejean, 1825: Bousquet (2012), Andrews (1923), Downie and Arnett (1996), MSU-ARC      

Clivina fossor fossor (Linnaeus, 1758): Bousquet (2012), Downie and Arnett (1996), MSU-ARC      

Clivina impressefrons LeConte, 1844: Bousquet (2012), Andrews (1923), Hatch (1924), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Clivina pallida Say, 1823: Bousquet (2012)      

Clivina rubicunda LeConte, 1857: Bousquet (2012)      

??Clivina rufa LeConte, 1857: Hubbard and Schwarz (1878); "The record from the lower peninsula of Michigan (Hubbard and Schwarz 1878: 644) needs confirmation." Bousquet (2012)

Paraclivina bipustulata (Fabricius, 1801): Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Paraclivina ferrea (LeConte, 1857): Petrillo and Witter (2009); reported from all surrounding states in Bousquet (2012)

Schizogenius amphibius (Haldeman, 1843): Bousquet (2012), Downie and Arnett (1996)      

Schizogenius ferrugineus Putzeys, 1846: Bousquet (2012), Hubbard and Schwarz (1878), MSU-ARC      

Schizogenius lineolatus (Say, 1823): Ruesink and Parsons (2022), MSU-ARC; ID of specimens in MSU-ARC by Peter W. Messer

Tribe Dyschiriini

Dyschirius aeneolus LeConte, 1850: Bousquet (2012), Leng (1920), Downie and Arnett (1996), Hubbard and Schwarz (1878)      

Dyschirius affinis Fall, 1901: Ruesink and Parsons (2022), MSU-ARC; ID of specimen in MSU-ARC by Peter W. Messer

Dyschirius brevispinus LeConte, 1878: Bousquet (2012), Leng (1920), Downie and Arnett (1996), Hubbard and Schwarz (1878)      

Dyschirius dejeanii Putzeys, 1846: Bousquet (2012), Dunn (1984), Hubbard and Schwarz (1878), MSU-ARC      

Dyschirius erythrocerus LeConte, 1857: Bousquet (2012), MSU-ARC      

Dyschirius globulosus (Say, 1823): Bousquet (2012), Dunn (1984), Leng (1920), Hubbard and Schwarz (1878), MSU-ARC      

Dyschirius haemorrhoidalis (Dejean, 1831): Bousquet (2012), MSU-ARC      

??Dyschirius longulus LeConte, 1850: Andrews (1923), Downie and Arnett (1996), Hubbard and Schwarz (1878); a northern sp., Bousquet (2012) says MI records are in error.

Dyschirius montanus LeConte, 1879: Bousquet (2012), MSU-ARC      

Dyschirius pallipennis (Say, 1823): Bousquet (2012), MSU-ARC      

Dyschirius pilosus LeConte, 1857: Bousquet (2012)      

Dyschirius politus politus (Dejean, 1825): Bousquet (2012), MSU-ARC      

Dyschirius sellatus LeConte, 1857: Messer (2021), Andrews (1923), MSU-ARC      

Dyschirius setosus LeConte, 1857: Bousquet (2012), Andrews (1923), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Dyschirius sphaericollis (Say, 1823): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Dyschirius terminatus LeConte, 1846: Bousquet (2012)      

Tribe Pasimachini

Pasimachus elongatus LeConte, 1846: Bousquet (2012), Townsend (1889)      

??Pasimachus sublaevis (Palisot de Beauvois, 1811): UCM:UCMC; ID needs verification, a bit north of expected range

Tribe Scaritini

Scarites subterraneus Fabricius, 1775: Bousquet (2012), Andrews (1916), BugGuide, Hubbard and Schwarz (1878), MSU-ARC      

Scarites vicinus Chaudoir, 1843: Ruesink and Parsons (2022), BugGuide, KY:UKIC , TAMU:TAMUIC , MSU-ARC; name resurrected from synonymy by Bousquet and Skelley (2010)

Subfamily Trechinae

Tribe Bembidiini

Amerizus wingatei (Bland, 1864): Bousquet (2012), Downie and Arnett (1996)      

Bembidion affine Say, 1823: Bousquet (2012), Dunn (1984), Andrews (1923), Downie and Arnett (1996), MSU-ARC      

Bembidion americanum Dejean, 1831: Bousquet (2012), Dunn (1984), Andrews (1916), Hubbard and Schwarz (1878), MSU-ARC      

Bembidion carinula Chaudoir, 1868: Bousquet (2012), Andrews (1923), BugGuide, Hubbard and Schwarz (1878), MSU-ARC      

Bembidion castor Lindroth, 1963: Bousquet (2012), MSU-ARC      

Bembidion chalceum Dejean, 1831: Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Bembidion cheyennense Casey, 1918: Bousquet (2012)      

Bembidion concolor (Kirby, 1837): Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Bembidion concretum Casey, 1918: Bousquet (2012), Leng (1920), Downie and Arnett (1996), MSU-ARC      

Bembidion confusum Hayward, 1897: Bousquet (2012), Leng (1920), Andrews (1923), Downie and Arnett (1996), MSU-ARC      

Bembidion cordatum (LeConte, 1847): Bousquet (2012), Dunn (1984), MSU-ARC      

Bembidion coxendix Say, 1823: Bousquet (2012)      

Bembidion dorsale Say, 1823: Bousquet (2012)      

Bembidion fortestriatum (Motschulsky, 1845): Bousquet (2012), Leng (1920), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Bembidion frontale (LeConte, 1847): Bousquet (2012), Andrews (1916), Hubbard and Schwarz (1878), MSU-ARC      

Bembidion fugax (LeConte, 1848): Bousquet (2012)      

Bembidion graciliforme Hayward, 1897: Bousquet (2012), MSU-ARC      

??Bembidion graphicum Casey, 1918: Downie and Arnett (1996); MI record doubtful, although could occur in western UP

Bembidion grapii Gyllenhal, 1827: Bousquet (2012), Adams (1909)      

??Bembidion honestum Say, 1823: Bousquet (2012), Hatch (1924); Bousquet (2012) says MI record needs confirmation

Bembidion impotens Casey, 1918: Bousquet (2012), Dunn (1984), Hatch (1924)      

Bembidion inaequale Say, 1823: Bousquet (2012), Andrews (1923), MSU-ARC      

Bembidion incrematum LeConte, 1860: Bousquet (2012), Leng (1920), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Bembidion intermedium (Kirby, 1837): Bousquet (2012), Hatch (1924), MSU-ARC      

Bembidion lacunarium (Zimmermann, 1869): Bousquet (2012), MSU-ARC      

Bembidion levettei carrianum Casey, 1924: Bousquet (2012), Hubbard and Schwarz (1878), MSU-ARC      

Bembidion levigatum Say, 1823: Bousquet (2012)      

Bembidion mimus Hayward, 1897: Bousquet (2012), Dunn (1984), MSU-ARC      

Bembidion muscicola Hayward, 1897: Bousquet (2012), Andrews (1923), Leng (1920), MSU-ARC      

Bembidion mutatum Gemminger & Harold, 1868: Bousquet (2012), Hubbard and Schwarz (1878), MSU-ARC      

Bembidion nigripes (Kirby, 1837): Bousquet (2012), Downie and Arnett (1996), MSU-ARC      

Bembidion nigrivestis Bousquet, 2006: Bousquet (2012)      

Bembidion nigrum Say, 1823: Bousquet (2012), Andrews (1916), MSU-ARC      

Bembidion nitidum (Kirby, 1837): Bousquet (2012), Andrews (1923), Hatch (1924), BugGuide, MSU-ARC      

Bembidion oberthueri Hayward, 1901: Bousquet (2012), MSU-ARC      

Bembidion obscurellum obscurellum (Motschulsky, 1845): Bousquet (2012), Dunn (1984), Andrews (1923), MSU-ARC      

Bembidion obtusum Audinet-Serville, 1821: Bousquet (2012), Purrington et al. (2000), MSU-ARC      

Bembidion occultator Notman, 1920: Bousquet (2012), MSU-ARC      

Bembidion patruele Dejean, 1831: Bousquet (2012), Dunn (1984), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Bembidion pedicellatum LeConte, 1857: Bousquet (2012), Hubbard and Schwarz (1878)      

Bembidion petrosum petrosum Gebler, 1833: Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Bembidion planatum (LeConte, 1847): Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Bembidion planum (Haldeman, 1843): Bousquet (2012)      

Bembidion praticola Lindroth, 1963: Bousquet (2012)      

Bembidion pseudocautum Lindroth, 1963: Bousquet (2012)      

Bembidion punctatostriatum Say, 1823: Bousquet (2012), MSU-ARC      

??Bembidion quadrimaculatum dubitans (LeConte, 1852): OSU:OSU; not known east of Manitoba

Bembidion quadrimaculatum oppositum Say, 1823: Bousquet (2012), Andrews (1923), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Bembidion rapidum (LeConte, 1847): Bousquet (2012), Dunn (1984), BugGuide, Downie and Arnett (1996), MSU-ARC      

Bembidion robusticolle Hayward, 1897: Bousquet (2012), Leng (1920), Downie and Arnett (1996)      

Bembidion rupicola (Kirby, 1837): Bousquet (2012)      

Bembidion salebratum (LeConte, 1847): Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Bembidion scopulinum (Kirby, 1837): Bousquet (2012), Andrews (1923), Hatch (1924), MSU-ARC      

Bembidion semicinctum Notman, 1919: Bousquet (2012), MSU-ARC      

Bembidion semistriatum (Haldeman, 1843): Bousquet (2012)      

Bembidion simplex Hayward, 1897: Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878)      

??Bembidion sordidum (Kirby, 1837): Bousquet (2012); Bousquet (2012) says MI record needs confirmation

Bembidion tetracolum tetracolum Say, 1823: Bousquet (2012), Andrews (1923), MSU-ARC      

??Bembidion timidum (LeConte, 1847): Bousquet (2012); Bousquet (2012) says MI record needs confirmation.

Bembidion transparens transparens (Gebler, 1830): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Bembidion transversale Dejean, 1831: Bousquet (2012), Dunn (1984), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Bembidion variegatum Say, 1823: Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Bembidion versicolor (LeConte, 1847): Bousquet (2012), Dunn (1984), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Bembidion versutum LeConte, 1878: Bousquet (2012), Downie and Arnett (1996), Hubbard and Schwarz (1878)      

Bembidion viridicolle (LaFerté-Sénectère, 1841): Ruesink and Parsons (2022), MSU-ARC      ID of MSU-ARC specimens by Peter W. Messer

Elaphropus anceps (LeConte, 1848): Bousquet (2012), MSU-ARC      

Elaphropus capax (LeConte, 1863): Ruesink and Parsons (2022), MSU-ARC; ID of specimen in MSU-ARC by Peter W. Messer

Elaphropus granarius (Dejean, 1831): Bousquet (2012)      

Elaphropus incurvus (Say, 1830): Bousquet (2012), Andrews (1923), Hatch (1924), Hubbard and Schwarz (1878), MSU-ARC      

Elaphropus tripunctatus (Say, 1830): Bousquet (2012), MSU-ARC      

Elaphropus vivax (LeConte, 1848): Bousquet (2012), Hubbard and Schwarz (1878)      

Elaphropus xanthopus (Dejean, 1831): Bousquet (2012), Hubbard and Schwarz (1878), MSU-ARC      

Micratopus aenescens (LeConte, 1848): Bousquet (2012)      

Mioptachys flavicauda (Say, 1823): Bousquet (2012), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC      

Paratachys proximus (Say, 1823): Ruesink and Parsons (2022), Hubbard and Schwarz (1878), MSU-ARC; ID of specimens in MSU-ARC by Peter W. Messer

Paratachys scitulus LeConte, 1848: Bousquet (2012), MSU-ARC      

Polyderis laeva (Say, 1823): Bousquet (2012), Hubbard and Schwarz (1878), MSU-ARC      

Porotachys bisulcatus (Nicolai, 1822): Bousquet (2012), MSU-ARC      

Tachyta angulata Casey, 1918: Bousquet (2012), Downie and Arnett (1996), MSU-ARC      

Tachyta inornata (Say, 1823): Bousquet (2012), Andrews (1923), Downie and Arnett (1996), Hubbard and Schwarz (1878), MSU-ARC      

Tribe Trechini

Blemus discus discus (Fabricius, 1792): Bousquet (2012), MSU-ARC      

Trechus apicalis Motschulsky, 1845: Bousquet (2012), Hubbard and Schwarz (1878), MSU-ARC      

Trechus quadristriatus (Schrank, 1781): Bousquet (2012)      

The following 43 species were reported from Michigan, but we consider them to be adventive, misidentifications or other errors:

Subfamily Brachininae

Tribe Brachinini

Brachinus conformis Dejean, 1831: Hubbard and Schwarz (1878), BYU:BYUC; a FL species, cannot be in MI

Brachinus explosus Erwin, 1970: BYU:BYUC; Mexico & AZ only, cannot be in MI

Brachinus lateralis Dejean, 1831: Townsend (1889); B. lateralis is a CA-AZ sp., cannot be in MI

Brachinus perplexus Dejean, 1831: Andrews (1916), Townsend (1889), BYU:BYUC; B. perplexus does not occur north of TN (Bousquet, 2012)

Brachinus puberulus  Chaudoir, 1868: BYU:BYUC; a Texas sp., cannot be in MI

Subfamily Carabinae

Tribe Carabini

Calosoma sycophanta (Linnaeus, 1758): Bousquet and Larochelle (1993), MSU-ARC      introduced for spongy moth control, no evidence that it is established in MI (Bousquet, 2012)

Subfamily Cicindelinae

Tribe Cicindelini

Cicindela hirticollis shelfordi Graves, 1988: CSU:CSUC; a western subspecies

Cicindela purpurea audubonii LeConte, 1845: CSU:CSUC; a western subspecies

Tribe Megacephalini

Tetracha carolina (Linneaus, 1763): UA:UAIC; a southern species, south of the Ohio River

Subfamily Elaphrinae

Tribe Elaphrini

Blethisa multipunctata (Linnaeus, 1758): Downie and Arnett (1996), Leng and Beutenmüller (1894b), Hubbard and Schwarz (1878); a palearctic sp. that does not occur in or near MI, see comment under Blethisa hudsonica.

Elaphrus laevigatus LeConte, 1852: Andrews (1923), Leng and Beutenmüller (1894b)      E. laevigatus is a CA sp.

Elaphrus riparius (Linnaeus, 1758): Andrews (1923), Hubbard and Schwarz (1878)      E. riparius is a Palearctic sp. not in NA.

Subfamily Harpalinae

Tribe Chlaeniini

Chlaenius circumcinctus Say, 1830: Townsend (1886), Townsend (1889); a southeastern sp.; cannot be in MI

Chlaenius harpalinus Eschscholtz, 1833: Hatch (1924); a western sp.; cannot be in MI

Tribe Galeritini

Galerita lecontei Dejean, 1831: Townsend (1889); a southern sp.; should not be in MI

Tribe Harpalini

Anisodactylus alternans (Motschulsky, 1845): CSU:CSUC as  Anisodactylus  nivalis; a western sp, prob. not in MI.

Anisodactylus rudis  LeConte, 1863: UA:UAIC; a far western sp., cannot be in MI

Harpalus amputatus  Say, 1830: UASM:UASM; a western sp., should not be in MI

Harpalus cautus Dejean, 1829: Hatch (1924); a western sp., should not be in MI

Stenolophus pallipes (Perroud & Montrouzier, 1864): KY:UKIC; not sure what this is, name appears in GBIF as valid for NA but not in any other standard source.

Tribe Lebiini

Microlestes nigrinus (Mannerheim, 1843): Adams (1909) as Blechrus nigrinus Mann.; a western sp., should not be in MI

Tribe Licinini

Dicaelus sculptilis sculptilis Say, 1823: Downie and Arnett (1996), Hubbard and Schwarz (1878); the closest this subsp. comes to MI is central Iowa.

Tribe Oodini

Oodes americanus Dejean, 1826: Andrews (1916); O. americanus is SE coastal, Andrews prob. had O. brevis Lindroth.

Tribe Platynini

Agonum limbatum Motschulsky, 1845: Andrews (1923) as Platynus variolatus Lec.; a western sp. that cannot be in MI

Agonum pallipes (Fabricius, 1787): Andrews (1916), Andrews (1923), BYU:BYUC as Platynus limbatus Say; a southern sp., should not be in MI

Platynus brunneomarginatus (Mannerheim, 1843): Andrews (1923), Hubbard and Schwarz (1878) as Platynus tenebricosus Gemm.; a far western sp., cannot be in MI

Platynus parmarginatus Hamilton, 1893: Andrews (1916); northern edge of range said to be near Ohio River

Sericoda bogemannii (Gyllenhal, 1813): Hatch (1924), Andrews (1916), Andrews (1923) as Agonum bogemanni (Gyll.); this western sp. should not occur in MI.

Tribe Pterostichini

Cyclotrachelus furtivus (LeConte, 1853): Wickham (1895), Leng (1920) as Eumolops furtiva (Lec.); Bousquet (2012) says MI record is probably in error, found E of Appalachians only

Cyclotrachelus  sigillatus (Say, 1823): iNaturalist Observations; an east coast & Appalachian sp., should not be in MI

Pterostichus pinguedineus (Eschscholtz, 1823): Hubbard and Schwarz (1878), Wolcott (1909) as Pterostichus mandibularis Kby.; a far northern sp., should not be in MI

Pterostichus superciliosus (Say, 1823): Wickham (1895); "The record from 'Michigan' (Wickham 1895: 185) is in error." Bousquet (2012), an eastern sp. that should not be in MI.

Pterostichus  oblongopunctatus  (Fabricius, 1787): UA:UAIC; European, cannot be in MI

Tribe Zabrini

Amara discors Kirby, 1837: Downie and Arnett (1996); a western sp., should not be in MI

Subfamily Nebriinae

Tribe Nebriini

Nebria sahlbergi Fischer von Waldheim, 1828: Andrews (1923); N. sahlbergi is western NA--not close to MI

Tribe Notiophilini

Notiophilus sibiricus Motschulsky, 1844: Hubbard and Schwarz (1878); a Palearctic sp. not in NA.

Subfamily Omophroninae

Tribe Omophronini

Omophron dentatum LeConte, 1852: Andrews (1916); a Calif. species.

Subfamily Scaritinae

Tribe Dyschiriini

Dyschirius edentulus Putzeys, 1846: Hubbard and Schwarz (1878); a TX-OK-FL sp. "The record from the lower peninsula of Michigan (Hubbard and Schwarz 1878: 644) is in error." Bousquet (2012)

Tribe Scaritini

Scarites quadriceps Chaudoir, 1843: UA:UAIC , Downie and Arnett (1996); not S. quadriceps, prob. refers to S. vicinus (Bousquet 2012)

Subfamily Trechinae

Tribe Bembidiini

Amerizus oblongulus (Mannerheim, 1852): Hubbard and Schwarz (1878); A. oblongulus occurs only on the Pacific coast. Bousquet (2012)

Bembidion anguliferum (LeConte, 1852): Andrews (1923), Hubbard and Schwarz (1878)      B. anguliferum is restricted to Calif.

Bembidion breve (Motschulsky, 1845): Hubbard and Schwarz (1878) as Bembidium tetraglyptum Mannh.; MSU-ARC has 4 specimens that appear to be Hubbard and Schwarz (1878) specimens from Marquette, they are actually B. occultator

Bembidion striola (LeConte, 1852): Hubbard and Schwarz (1878); a west coast sp., cannot be in MI.

This family list was last updated: April 15, 2022