Introduction. The name of each listed species is followed by up to five sources that report the species as occurring in Michigan. Sources in all upper-case letters refer to specific insect collections: MSU-ARC refers to material held by the Michigan State University Arthropod Research Collection (data provided by G. Parsons). All others refer to collection data extracted from the Symbiota Collections of Arthropods Network (SCAN) database online at https://scan-bugs.org/. See the Methods page for a list of abbreviations and associated collections.
Species names preceded by a single '?' are considered probable (for occurring in Michigan) but uncertain records. Those preceded by '??' are considered possible but unlikely to occur in Michigan. Species that are fully accepted as having valid Michigan records are unmarked. Species reported in the literature that we consider to be erroneous Michigan records are in a separate list following the others.
For Deinopsis americana some sources use () on Kraatz and others do not. Apparently Agassiz emended Deinopsis to Dinopsis in 1846, and Kraatz described americanus in Dinopsis. We do not know the rules well enough to know whether () are required in this case.
STAPHYLINIDAE - ALEOCHARINAE
1 subfamily, 13 tribes, 39 genera, 81 species
Tribe Aleocharini
Aleochara bimaculata Gravenhorst, 1802: Klimaszewski (1984), Andrews (1923), Hatch (1925), Downie and Arnett (1996), MSU-ARC
Aleochara curtula (Goeze, 1777): Pastula and Merritt (2013), Hubbard and Schwarz (1878), MSU-ARC
Aleochara gracilicornis Bernhauer, 1901: Klimaszewski (1984), Downie and Arnett (1996), MSU-ARC
Aleochara inexpectata Klimaszewski, 1984: Klimaszewski (1984), Downie and Arnett (1996)
Aleochara lanuginosa Gravenhorst, 1802: Ruesink and Parsons (2024), MSU-ARC
Aleochara lata Gravenhorst, 1802: Klimaszewski (1984), Andrews (1923), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC
Aleochara lustrica Say, 1832: Klimaszewski (1984), Downie and Arnett (1996)
?Tinotus morion Gravenhorst, 1802: Hatch (1925); a European sp., widespread but uncommon in NA
??Aleochara ocularis Klimaszewski, 1984: Downie and Arnett (1996); where did their info come from?
Aleochara sculptiventris (Casey, 1893): Hatch (1925), Ruesink and Parsons (2024), MSU-ARC
Aleochara speculicollis Bernhauer, 1901: Klimaszewski (1984), Downie and Arnett (1996)
Aleochara thoracica Casey, 1893: Klimaszewski (1984), Downie and Arnett (1996), MSU-ARC
Aleochara verna Say, 1833: Klimaszewski (1984), Downie and Arnett (1996), MSU-ARC
Tribe Athetini
??Acrotona recondita (Erichson, 1839): Hatch (1925) as Arisota recondita Er.
Amischa analis (Gravenhorst, 1802): Klimaszewski (2018b), Hubbard and Schwarz (1878), MSU-ARC
Atheta aemula (Erichson, 1839): Ruesink and Parsons (2024), Andrews (1923), MSU-ARC; not common but widespread in eastern N.A.
Atheta festinans (Erichson, 1839): Klimaszewski et al. (2018b), Downie and Arnett (1996), Andrews (1923), Leng (1920), Townsend (1889)
Atheta modesta (Melsheimer, 1844): Gusarov (2003), Hatch (1925), Klimaszewski et al. (2018a)
Meronera venustula (Erichson, 1839): Klimaszewski et al. (2018b), Hubbard and Schwarz (1878), MSU-ARC
Mocyta luteola (Erichson, 1839): Klimaszewski et al. (2018a), Bernhauer (1907), Downie and Arnett (1996)
Nehemitropia lividipennis (Mannerheim, 1830): Klimaszewski et al. (2018b), Andrews (1923), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC
?Philhygra wisconsinica (Casey, 1910): Hatch (1925); GBIF places it in the genus Atheta
Sableta infulata Casey, 1910: Ashe and Gusarov (2004)
Seeversiella globicollis (Bernhauer, 1907): Ashe (1986), Klimaszewski et al. (2018a), Newton et al. (2001)
Tribe Autaliini
Autalia rivularis (Gravenhorst, 1802): Klimaszewski (2018b), Hoebeke (1988), MSU-ARC
Tribe Falagriini
Aleodorus bilobatus Say, 1833: Klimaszewski et al. (2018b), Casey (1906), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC
Aleodorus scutellaris (LeConte, 1866): Klimaszewski et al. (2018b), Downie and Arnett (1996)
Cordalia obscura (Gravenhorst, 1802): Klimaszewski et al. (2018b), Downie and Arnett (1996), MSU-ARC
Falagria dissecta Erichson, 1839: Klimaszewski et al. (2018a), Hatch (1925), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC
Myrmecocephalus cingulatus (LeConte, 1866): Klimaszewski et al. (2018b), Hubbard and Schwarz (1878), Downie and Arnett (1996)
?Myrmecopora vaga (LeConte, 1866): Downie and Arnett (1996); location MI (Lake Superior)
Tribe Gymnusini
Adinopsis bicornis Klimaszewski, 1979: Klimaszewski (1982), Klimaszewski and Frank (1992b), Downie and Arnett (1996), MSU-ARC
Deinopsis harringtoni Casey, 1911: Klimaszewski (1982), Klimaszewski and Frank (1992b), Downie and Arnett (1996)
Deinopsis illinoisensis Klimaszewski, 1979: Klimaszewski and Frank (1992b), Downie and Arnett (1996)
Gymnusa atra Casey, 1911: Klimaszewski et al. (2018a), MSU-ARC
Gymnusa grandiceps Casey, 1915: Klimaszewski and Frank (1992b), MSU-ARC
Gymnusa pseudovariegata Klimaszewski, 1979: Ruesink and Parsons (2024), MSU-ARC
Tribe Homalotini
Eumicrota corruscula (Erichson, 1839): Klimaszewski et al. (2009), Hubbard and Schwarz (1878), Seevers (1951), MSU-ARC
Eumicrota socia (Erichson, 1839): Klimaszewski et al. (2009), Hubbard and Schwarz (1878), Seevers (1951), Downie and Arnett (1996)
Gyrophaena affinis Sahlberg, 1834: Klimaszewski et al. (2009), Seevers (1951), MSU-ARC
Gyrophaena blatchleyi Seevers, 1951: Enushchenko (2018), Seevers (1951), Downie and Arnett (1996), MSU-ARC
Gyrophaena caseyi Seevers, 1951: Klimaszewski et al. (2009), Seevers (1951), Downie and Arnett (1996)
Gyrophaena chippewa Seevers, 1951: Klimaszewski et al. (2009), Seevers (1951), Downie and Arnett (1996)
Gyrophaena coniciventris Casey, 1906: Seevers (1951), MSU-ARC
Gyrophaena flavicornis Melsheimer, 1844: Klimaszewski et al. (2009), Seevers (1951), Hubbard and Schwarz (1878), MSU-ARC
Gyrophaena gaudens Casey, 1906: Klimaszewski et al. (2009), Seevers (1951), Downie and Arnett (1996), MSU-ARC
Gyrophaena glivicollis Casey, 1906: Klimaszewski et al. (2009), Seevers (1951), MSU-ARC
Gyrophaena hubbardi Seevers, 1951: Seevers (1951), Downie and Arnett (1996); paratypes incl. a Detroit specimen
Gyrophaena insolens Casey, 1906: Klimaszewski et al. (2009), Seevers (1951), Downie and Arnett (1996)
Gyrophaena lobata Casey, 1906: Klimaszewski et al. (2009), Seevers (1951), MSU-ARC
Gyrophaena michigana Seevers, 1951: Seevers (1951), Downie and Arnett (1996)
Gyrophaena modesta Casey, 1906: Klimaszewski et al. (2009), Seevers (1951), Majka and Klimaszewski (2011), MSU-ARC
Gyrophaena nana (Paykull, 1800): Klimaszewski et al. (2009), Seevers (1951), Hatch (1925), Downie and Arnett (1996), MSU-ARC
?Gyrophaena simulans Seevers, 1951: FMNH:FMNHINS; Benzie Co.
Gyrophaena subnitens Casey, 1906: Majka and Klimaszewski (2011), Klimaszewski et al. (2009), Seevers (1951), Downie and Arnett (1996)
Gyrophaena vitrina Casey, 1906: Klimaszewski et al. (2009), Seevers (1951), MSU-ARC
?Gyrophaena wisconsinica Seevers, 1951: FMNH:FMNHINS; Berrien Co.
Phanerota dissimilis (Erichson, 1839): Enushchenko (2018), Seevers (1951), Hubbard and Schwarz (1878), MSU-ARC
Phanerota fasciata (Say, 1832): Klimaszewski (2018b), Seevers (1951), Hubbard and Schwarz (1878), MSU-ARC
??Phymatura blanchardi(Casey, 1893): KU:SEMC; Eaton Co, Biological Reserve
Pleurotobia trimaculata (Erichson, 1839): Andrews (1923), Hubbard and Schwarz (1878), Ruesink and Parsons (2024), MSU-ARC
Silusida marginella (Casey, 1893): Hatch (1925), Ruesink and Parsons (2024), MSU-ARC
Tribe Hoplandriini
Hoplandria lateralis (Melsheimer, 1844): Genier (1989), Andrews (1923), Hubbard and Schwarz (1878), MSU-ARC
Tribe Hypocyphtini
??Oligota pumilio Kiesenwetter, 1858: Schwarz (1876), Hubbard and Schwarz (1878) as Oligota pedalis Lec.
Tribe Lomechusini
Myrmoecia lauta (Casey, 1893): Klimaszewski et al. (2005), Downie and Arnett (1996), MSU-ARC
Xenodusa cava (LeConte, 1863): Klimaszewski et al. (2018b), Hubbard and Schwarz (1878), Downie and Arnett (1996), MSU-ARC
Xenodusa reflexa (Walker, 1866): Klimaszewski et al. (2018a), Downie and Arnett (1996), MSU-ARC
Zyras obliquus (Casey, 1893): Klimaszewski et al. (2018a)
Tribe Myllaenini
Myllaena insomnis Casey, 1911: Klimaszewski and Frank (1992a)
Myllaena ludificans Casey, 1911: Klimaszewski et al. (2018b)
??Myllaena vulpina Bernhauer, 1907: Hatch (1925); known from ON, PA, KY & AR; MI record needs confirmation.
Tribe Oxypodini
Blepharhymenus lacustris (Casey, 1893): Casey (1893), Downie and Arnett (1996), SI:NMNH; described from a MI specimen
Devia prospera (Erichson, 1839): Gusarov (2003), Klimaszewski et al. (2018a), Downie and Arnett (1996), MSU-ARC
??Oxypoda nigriceps Casey, 1893: Hatch (1925)
?Oxypoda vockerothi Klimaszewski, 2006: Ruesink and Parsons (2024), MSU-ARC
Tribe Placusini
Euvira micmac Klimaszewski and Majka, 2007: Klimaszewski and Majka (2007)
Placusa canadensis Klimaszewski, 2001: Klimaszewski et al. (2001)
Placusa despecta Erichson, 1839: Klimaszewski et al. (2001), Townsend (1889), Peck and Thomas (1998)
Tribe Tachyusini
?Gnypeta nigrella (LeConte, 1863): Hatch (1925), MSU-ARC; IN in Downie and Arnett (1996) and IL & ON in Klimaszewski et al. (2018a), we cannot confirm that MSU specimens are correctlt IDed
Tachyusa americana Casey, 1906: Ruesink and Parsons (2024), MSU-ARC
Tachyusa gracillima LeConte, 1863: Ruesink and Parsons (2024), MSU-ARC; or this might be T. smetanai Pasnik, 2006. We are uncertain which of these two species we have, but since neither has been reported from Michigan, we are including this record here.
The following species were also reported from Michigan, but we consider them to be adventive, misidentifications or other errors:
Tribe Aleocharini
Aleochara bipustulata (Linnaeus, 1760): Hatch (1925), Hubbard and Schwarz (1878) as Aleochara nitida Grav.; A. bipustulata is a palearctic sp., does not occur in NA
Tribe Athetini
Atheta divisa (Märkel, 1844): Hatch (1925); a European sp., does not occur in NA
Mocyta fungi (Gravenhorst, 1806): Hatch (1925); a European sp., does not occur in NA
Tribe Gymnusini
Deinopsis americana (Kraatz, 1857): Schwarz (1876), Hubbard and Schwarz (1878) as Dinopsis myllaenoides Kr.; a southern sp.
Gymnusa brevicollis (Paykull, 1800): Hubbard and Schwarz (1878), Leng (1920); a European sp., does not occur in NA
Gymnusa variegata Kiesenwetter, 1845: Hubbard and Schwarz (1878), Leng (1920); a European sp., does not occur in NA
Tribe Gyrophaenini
Sternotropa elevata (Fauvel, 1889): Townsend (1889) as Homalota elevata Fauv.; a sp. from New Caledonia (as Brachida elevata), not known to occur in NA.
Tribe Homalotini
Homalota plana (Gyllenhal, 1810): Townsend (1889), Andrews (1923); a Eurasian sp. thought to have arrived in America much later than 1923
Tribe Mesoporini
Anacyptus testaceus (LeConte, 1863): Peck and Thomas (1998), Downie and Arnett (1996); Michigan record as MI (Lake Superior), but supposedly a southern species
Tribe Myllaenini
Myllaena dubia (Gravenhorst, 1806): Hubbard and Schwarz (1878); a European sp., does not occur in NA
Myllaena minuta (Gravenhorst, 1806): Schwarz (1876), Hubbard and Schwarz (1878) as Myllaena fuscipennis Kr.; a European sp., does not occur in NA
Tribe Oxypodini
Crataraea suturalis (Mannerheim, 1830): Townsend (1889) as Microglossa suturalis Mann.; a European sp., does not occur in NA
Phloeopora testacea (Mannerheim, 1830): Hatch (1925); a European sp., does not occur in NA
This family list was last updated: April 7, 2023